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Here - The Alfred Russel Wallace Website

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16 THE WORLD OF LIFE CHAP<br />

Another point of some importance is the greater stability<br />

in the flora of meadow as compared with that of pasture<br />

land. In the former only one plant was an accidental<br />

straggler, while in the latter there were 1 2, or two-thirds of<br />

the peculiar species. <strong>The</strong>se are mostly rare, and are very often<br />

not truly British plants, so that they cannot be considered<br />

as permanent pasture plants. <strong>The</strong> more stable meadow<br />

flora is no doubt largely due to the fact that few of the late-<br />

flowering plants are allowed to produce seed, and though<br />

seed may be often introduced by birds or the wind, many<br />

of these species soon die out It thus appears that though<br />

pastures are actually richer in species than meadows, yet the<br />

latter have a more permanent character, as almost all those<br />

peculiar to pastures are comparatively rare and therefore<br />

very liable to disappear through very slight changes of<br />

conditions.<br />

<strong>The</strong>se various<br />

here given, serve<br />

facts, and many others<br />

to show us how very<br />

which cannot be<br />

delicate are the<br />

mutual relations and adjustments of plants to their total<br />

environment. In proportion as that environment is subject<br />

to change of any kind, some rare species die out, while others<br />

become diminished in numbers. And what takes place in<br />

single fields or other small areas, when closely studied, must<br />

certainly occur on a much grander scale over the whole<br />

earth, and especially in those countries and periods when<br />

great changes of climate or of physical geography are taking<br />

place. <strong>The</strong>se detailed studies of " Meadow and Pasture<br />

Analysis" as their author terms them thus demonstrate<br />

on a very small scale that " struggle for existence " which, as<br />

we shall see further on, is always present, acts in an almost<br />

infinite number of ways, and is one of the most important<br />

factors in the developmental changes of the World of Life.<br />

We will now proceed to give some of the numerical facts of<br />

plant distribution, in various areas small and large,<br />

as well<br />

as over the whole earth ; but it will be advisable first to<br />

give a brief account of the way in which this is usually<br />

dealt with by botanists.<br />

Four years before the appearance of the Origin of<br />

Species the great Swiss botanist, Alphonse De Candolle<br />

published one of the most remarkable and interesting

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