Sheep - AgRIS

Sheep - AgRIS Sheep - AgRIS

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very mediocre parents and ancestors. For breeding purposes, it would be much more desirable to keep superior individuals from parents and ancestors that themselves were outstanding. b) Pedigrees A pedigree is a record of an individual ‗s a,ncestors that are related to him through his parents. Earlier, the information included in a pedigree has been simply the names and registration numbers of the ancestors, and little has been indicated as to the type and performance of the ancestors. Pedigrees now m include information on the size of the litter at birth and weaning. It full information is available on the ancestors as well as the collateral relatives, may be of importance in detecting carriers of a recessive gene. Such information has been used to a great extent in combating dwarfism in beef cattle. A disadvantage of the use of the pedigree in selection against a recessive gene is that there are often unintentional and unknown mistakes in pedigrees that may result in the condemnation of an entire line of breeding when actually the family may be free of such a defect. On the other hand, the frequency of a recessive gene in a family may be very low, and records may be incomplete. Then, later, it will be found that the gene is present, Another disadvantage of pedigree selection is that the individuals in the pedigree, especially the males, may have been selected from a very large group, and the pedigree tells us nothing about the merit of their relatives. Still another disadvantage of pedigree selection is that a pedigree may often become popular because of fashion or fad and not because of the merit of the individuals it contains. The popularity of the pedigree may change in a year or two, and the value of such a pedigree may decrease considerably or may even be discriminated against. If popularity is actually based on merit, there is less danger of a diminution of value in a short period of time. In using pedigrees for selection purposes, weight should be given to the most recent ancestors. This is because the percentage of genes contributed by an individual‘s ancestors is halved each new generation. Some breeders place much emphasis on some outstanding ancestor three or four generations removed in the pedigree, but such an ancestor contributes a very small percentage of the genes the individual possesses and has very little influence on type and performance, unless linebreeding to that ancestor has been practised. An individuals own performance is usually of more value in selection than its pedigree, but the pedigree may be used as an accessory to sway the balance when two animals are very similar in individuality but one has a more desirable pedigree than the other. Pedigree information is also quite useful whenthe animals are selected at a young age and their own type and conformation is not known. Pedigree is useful in identifying superior families if good records are kept and are available. c) Collateral Relatives Collateral relatives are those that are not related directly to an individual, either as ancestors or as their progeny. Thus, they are the individuals brothers, sisters, cousions, uncles and aunts. The more closely they are related to the individual in question, the more valuable is the information for selection purposes. Complete information on collateral relatives, gives an idea of the kinds of genes and combinations of genes that the individual is likely to possess. Information of this kind has been used in meathog certification programs, where a barrow and a gilt from each litter may be slaughtered to obtain carcass data. This is done, because otherwise the animal himself has to be slaughtered if information on his own carcass quality is to be obtaineddlnformation on collateral relatives is also used in selecting dairy bulls, since miLk production can be measured only in the cows even though the bull transmits genes to his offspring for this trait. Information on collaterals has been used in the All India coordinated Research on breeding Sheep for mutton where in information on slaughter traits has been used from full brothers slaughtered for selecting rams for future breeding. d) Progeny Test 381

Selection on this basis means that we estimate the breeding value of an individual through a study of the traits or characteristics of its offspring. In other words, the progeny of different individuals are studied to determine which group is superior, and on this basisthe superior breedingdms given preference for future breeding purposes. If information is complete, this is an excellent way of identifying superior breeding animals. Progeny tests are very useful for determining characteristics that are expressed only in one sex, such as milk production in cows or egg production in hens. Even though the bull does not produce milk nor does the rooster lay eggs, they carry genes for these traits and supply one-half of the inheritance of each of their daughters for that particular trait. Progeny tests are also useful in measuring traits which cannot be measured in the living individual. A good example ofthis is carcass quality in cattle, sheep, and hogs. Progeny tests are also being used at the present time by experiment stations in studies of reciprocal recurrent selection. This type of selection is used to test for the ―nicking, ability‖ of individuals and lines and is based on the performance of the line-cross progeny. Selection of this type is for traits that are lowly heritable and in which nonadditive gene action seems to be important. In comparing individuals on the basis of their progeny, certain precautions should be taken to make the comparisons fair and accurate. In conducting a progeny test, it is very important to test a random sample of the progeny. It would be more desirable if all progeny could be tested, but where this cannot be done, as in litters of swine, those nearer the average of the litter should be tested. It is also important that the females to which a male is mated should be a nonselected group. One would expect the offspring of a sire to be superior if he is mated to the outstanding females in the herd. Such a practice would be misleading in comparing males by a progeny test, since much of the superiority of the offspring of one male could come from the dams and not from the sire. Some breeders prefer using a rotation of different dams when testing males, but this is practical only in swine, where two litters may be produced each year. Using a large number of offspring in testing a sire increases the accuracy of the test rule. Where the number of females in a herd is limited, the number of males that may be progenytested will be less as the number of matings per sire is increased. The point is, then, that the breeder must make some decision as to how many sires to test and how many progeny must be produced to give a good test. The number of offspring required for an accurate progeny test will depend upon the heritability of a trait, with fewer offspring being required, when the trait is highly heritable, and more being required when it is lowly heritable. To make accurate progeny tests, it is also important to keep the environment as nearly as possible the same for the offspring of the different sires. In progeny-testing swine, for instance, confusion would result when the progeny of one sire were fed in dry lot during the summer and the progeny of another were fed on pasture. This would be particularly true in progeny-testing for rate of gain, where pigs fed modern rations often grow considerably faster in dry lot than on pasture. when this environmental condition were not controlled, the inferior sire might actually be thought to be superior. Progeny tests in most of our farm animals have certain definite limitations. In cattle, especially it takes so long to prove an animal on a progeny test that he may be dead before the test is completed and his merit actually known. Progeny tests may be tnow easily in swine than in other farm animals, but even in this case the males are usually disposed of by the time they are thoroughly progenytested. The process of progeny-testing may be speeded up by testing males at an earlier age than they~would ordinarily be used for breeding purposes. By hand-mating them to a few females, or by using them on a larger number of females by artificial insemination, harmful effects that might occur from overuse at too early an age may be prevented. Too often, farmers send their old sires to market just as soon as their daughters are old enough to breed, in order to prevent inbreeding. This practice has resulted in much loss of good genetic material for livestock improvement. Actually a sire is not proved until his daughters come into production. Rather than being slaughtered, a sire that has proved himself to be of high genetic merit should be used more extensively. It is true that his usefulness in a particular herd may be finished when his daughters are of breeding age, but he should be sent to another 382

Selection on this basis means that we estimate the breeding value of an individual through<br />

a study of the traits or characteristics of its offspring. In other words, the progeny of different<br />

individuals are studied to determine which group is superior, and on this basisthe superior<br />

breedingdms given preference for future breeding purposes. If information is complete, this is<br />

an excellent way of identifying superior breeding animals.<br />

Progeny tests are very useful for determining characteristics that are expressed only in one<br />

sex, such as milk production in cows or egg production in hens. Even though the bull does not<br />

produce milk nor does the rooster lay eggs, they carry genes for these traits and supply one-half<br />

of the inheritance of each of their daughters for that particular trait.<br />

Progeny tests are also useful in measuring traits which cannot be measured in the living<br />

individual. A good example ofthis is carcass quality in cattle, sheep, and hogs.<br />

Progeny tests are also being used at the present time by experiment stations in studies of<br />

reciprocal recurrent selection. This type of selection is used to test for the ―nicking, ability‖ of<br />

individuals and lines and is based on the performance of the line-cross progeny. Selection of<br />

this type is for traits that are lowly heritable and in which nonadditive gene action seems to be<br />

important.<br />

In comparing individuals on the basis of their progeny, certain precautions should be taken<br />

to make the comparisons fair and accurate. In conducting a progeny test, it is very important to<br />

test a random sample of the progeny. It would be more desirable if all progeny could be tested,<br />

but where this cannot be done, as in litters of swine, those nearer the average of the litter should<br />

be tested. It is also important that the females to which a male is mated should be a nonselected<br />

group. One would expect the offspring of a sire to be superior if he is mated to the outstanding<br />

females in the herd. Such a practice would be misleading in comparing males by a progeny test,<br />

since much of the superiority of the offspring of one male could come from the dams and not<br />

from the sire. Some breeders prefer using a rotation of different dams when testing males, but<br />

this is practical only in swine, where two litters may be produced each year.<br />

Using a large number of offspring in testing a sire increases the accuracy of the test rule.<br />

Where the number of females in a herd is limited, the number of males that may be progenytested<br />

will be less as the number of matings per sire is increased. The point is, then, that the<br />

breeder must make some decision as to how many sires to test and how many progeny must be<br />

produced to give a good test. The number of offspring required for an accurate progeny test will<br />

depend upon the heritability of a trait, with fewer offspring being required, when the trait is<br />

highly heritable, and more being required when it is lowly heritable.<br />

To make accurate progeny tests, it is also important to keep the environment as nearly as<br />

possible the same for the offspring of the different sires. In progeny-testing swine, for instance,<br />

confusion would result when the progeny of one sire were fed in dry lot during the summer and<br />

the progeny of another were fed on pasture. This would be particularly true in progeny-testing<br />

for rate of gain, where pigs fed modern rations often grow considerably faster in dry lot than on<br />

pasture. when this environmental condition were not controlled, the inferior sire might actually<br />

be thought to be superior.<br />

Progeny tests in most of our farm animals have certain definite limitations. In cattle,<br />

especially it takes so long to prove an animal on a progeny test that he may be dead before the<br />

test is completed and his merit actually known. Progeny tests may be tnow easily in swine than<br />

in other farm animals, but even in this case the males are usually disposed of by the time they<br />

are thoroughly progenytested.<br />

The process of progeny-testing may be speeded up by testing males at an earlier age than<br />

they~would ordinarily be used for breeding purposes. By hand-mating them to a few females,<br />

or by using them on a larger number of females by artificial insemination, harmful effects that<br />

might occur from overuse at too early an age may be prevented.<br />

Too often, farmers send their old sires to market just as soon as their daughters are old<br />

enough to breed, in order to prevent inbreeding. This practice has resulted in much loss of good<br />

genetic material for livestock improvement. Actually a sire is not proved until his daughters<br />

come into production. Rather than being slaughtered, a sire that has proved himself to be of<br />

high genetic merit should be used more extensively. It is true that his usefulness in a particular<br />

herd may be finished when his daughters are of breeding age, but he should be sent to another<br />

382

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