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A review of dipterocarps - Center for International Forestry Research

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Seed Handling 79<br />

Table 2. (continued) Seed (fruit) weight and size indicators at harvest.<br />

Species Mean seeds per kilo Mean length (mm) Mean width (mm)<br />

Dryobalanops rappa 1400 17 9<br />

Shorea faguetiana 1400 n/a n/a<br />

Shorea laevis 1600 14 9<br />

Anisoptera marginata 1800 10 10<br />

Shorea leprosula 1800 16 10<br />

Shorea affinis 1900 n/a n/a<br />

Shorea leptoderma 1900 n/a n/a<br />

Shorea ovata 1900 n/a n/a<br />

Dipterocarpus intricatus 2800 20 17<br />

Cotylelobium burckii 2900 10 10<br />

Cotylelobium melanoxylon 2900 9 8<br />

Shorea obtusa 2900 n/a n/a<br />

Hopea dryobalanoides 4000 10 7<br />

Vatica odorata ssp. odorata 4000 8 7<br />

Hopea parviflora 4100 7 6<br />

Shorea guiso 4200 11 6<br />

Hopea odorata 5300 8 6<br />

Hopea foxworthyi 5500 8 5<br />

Hopea ferrea 5800 n/a n/a<br />

Hopea mengerawan 6300 10 4<br />

Hopea nigra 9000 8 5<br />

Vatica mangachapoi 17000 5 5<br />

Monotes kerstingii* 45000 n/a n/a<br />

*: Assessment refers to seeds inside the fruit.<br />

to germinate they can be saved by storing in rigid<br />

containers lined with moist newspaper or other absorbent<br />

materials to keep them moist during transit.<br />

Size Considerations<br />

There is a large range in sizes <strong>of</strong> dipterocarp seeds (Table<br />

2) which implies that different handling procedures may<br />

be needed <strong>for</strong> moist seed <strong>of</strong> particular size ranges. For<br />

example, smaller seeds (< 2 g) would benefit from the<br />

inclusion <strong>of</strong> packing material to increase the size <strong>of</strong> air<br />

spaces between the seeds. Crumpled newspapers and<br />

polystyrene chips have been used <strong>for</strong> this purpose.<br />

Seed Processing<br />

The fruit <strong>of</strong> dipterocarp species, which is the unit<br />

employed <strong>for</strong> handling, is generally referred to as the<br />

‘seed’. Removal <strong>of</strong> calyx lobes (‘wings’) by manual<br />

abscission is carried out <strong>for</strong> all physiology types. This<br />

enables easier sowing and better contact <strong>of</strong> the seeds with<br />

the soil.<br />

Factors which should be considered in the drying <strong>of</strong><br />

OLDA seeds <strong>for</strong> storage are described in the summary<br />

at the end <strong>of</strong> the chapter.<br />

Insect infestation can be a major problem in the<br />

handling <strong>of</strong> seed, especially in the genus Dipterocarpus<br />

(Table 3, Prasad and Jalil 1987, Eungwijarnpanya and<br />

Hedlin 1984); sometimes 100% <strong>of</strong> individual seedlots are<br />

rendered useless. Methods to reduce this problem are<br />

required and would be best supported by studies on insect<br />

behaviour. Some studies have already been carried out<br />

on recalcitrant material <strong>of</strong> the rain <strong>for</strong>est (Toy et al.<br />

1992, Toy and Toy 1992); extension <strong>of</strong> such studies to<br />

include seasonal-climate species would be advantageous.<br />

Further discussion <strong>of</strong> infestation problems can be found<br />

in Chapter 7.<br />

Methods <strong>for</strong> Storage <strong>of</strong> Dipterocarp<br />

Seeds<br />

In the past half century, various methods <strong>of</strong> storage have<br />

been proposed <strong>for</strong> recalcitrant dipterocarp seeds and,

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