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A review of dipterocarps - Center for International Forestry Research

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Seed Physiology 69<br />

Dipterocarp seed <strong>of</strong> the OLDA type has a shorter<br />

storage life than seed <strong>of</strong> crop species if compared at the<br />

same moisture content. Thus, it has been estimated that<br />

the relevant K E and, C W viability constants (which<br />

indicate seed longevity) are, respectively, only 6.4 and<br />

2.9 on the average <strong>for</strong> dipterocarp seed (Table 6),<br />

compared with 8.4 and 4.7 (Tompsett 1994) <strong>for</strong><br />

herbaceous crops. Further research is needed to extend<br />

these findings to other OLDA dipterocarp species.<br />

The stage <strong>of</strong> fruit development at harvest is important<br />

to ensure optimum desiccation tolerance, and<br />

consequently to ensure maximum storage potential.<br />

Further research is needed <strong>for</strong> <strong>dipterocarps</strong> in order to<br />

closely assess the relationship between harvest<br />

condition, postharvest handling, and desiccation<br />

tolerance.<br />

Studies are needed to increase knowledge <strong>of</strong> the<br />

optimum moisture and temperature conditions <strong>for</strong><br />

storage <strong>of</strong> recalcitrant seeds, employing controlled<br />

conditions. Especially, research is needed in relation to<br />

the chilling injury. Studies to quantify chilling damage<br />

in relation to moisture content are needed. Also, research<br />

is required to determine its relationship to underlying<br />

biochemical processes.<br />

Although the database DABATTS (Tompsett and<br />

Kemp 1996a, b) includes a large amount <strong>of</strong> previously<br />

unpublished in<strong>for</strong>mation on dipterocarp seed, a high<br />

proportion <strong>of</strong> its contents are the results produced by<br />

the authors. Unpublished in<strong>for</strong>mation from other sources<br />

needs to be databased, building on DABATTS and<br />

increasing the total sum <strong>of</strong> research results readily<br />

available.<br />

<strong>Research</strong> on the induction <strong>of</strong> flowering is necessary<br />

to improve knowledge <strong>of</strong> the causes underlying the<br />

irregular flowering <strong>of</strong> <strong>dipterocarps</strong>. Such research may<br />

provide artificial means <strong>for</strong> the induction <strong>of</strong> flowering<br />

in relation to breeding and to seed production in nonmast<br />

years.<br />

These approaches might with benefit be extended to<br />

other tropical tree families such as the Palmae and<br />

Sapotaceae.<br />

Relevant Institutions<br />

As described in Chapter 4 and above (in the work <strong>of</strong><br />

Sasaki, Mori, Tang, Tamari and Yap), Forest <strong>Research</strong><br />

Institute Malaysia (FRIM) has played a leading role in<br />

early dipterocarp seed research, particularly in the areas<br />

<strong>of</strong> germination ecology and storage research. Current<br />

work at FRIM on cryopreservation and seedling storage<br />

is referred to elsewhere. Over the last decade, the seed<br />

physiological studies at the Royal Botanic Gardens Kew<br />

have contributed basic knowledge, creating a firm<br />

foundation <strong>for</strong> practical recommendations. The Forest<br />

<strong>Research</strong> Centre, Sandakan, Malaysia, has a seed research<br />

laboratory constructed under an FAO aid programme and<br />

has undertaken significant dipterocarp research.<br />

In Thailand the Royal Forest Department’s ASEAN<br />

Tree Seed Centre, Muak Lek, has been involved in<br />

dipterocarp studies <strong>for</strong> a number <strong>of</strong> years and has good<br />

facilities; additionally, the central laboratory in Bangkok<br />

has an active research team on the topic.<br />

The Ecosystems <strong>Research</strong> and Development Bureau,<br />

Republic <strong>of</strong> the Philippines, is engaged in dipterocarp<br />

seed research, as are the Forest <strong>Research</strong> and<br />

Development Centre, the Biotechnology Centre and<br />

BIOTROP in Bogor, Indonesia. In India, research on<br />

biochemical aspects has been recently conducted at the<br />

High Altitude Plant Physiology <strong>Research</strong> Centre <strong>of</strong><br />

Garhwal University, Srinagar and the Forest <strong>Research</strong><br />

Institute, Dehra Dun has been involved in dipterocarp<br />

research in the recent past.<br />

In China, biochemical, ultrastuctural and<br />

physiological research on dipterocarp species has been<br />

per<strong>for</strong>med by staff <strong>of</strong> the Tropical Forest <strong>Research</strong><br />

Institute, Chinese Academy <strong>of</strong> <strong>Forestry</strong>, Guangdong.<br />

Although not involved in dipterocarp research, the<br />

Agriculture and Horticulture Department at Reading<br />

University, UK, is developing experience in the area <strong>of</strong><br />

tropical tree seed physiology. Other institutes have<br />

contributed in<strong>for</strong>mation in this field, but space available<br />

limits the numbers that can be included.<br />

Acknowledgements<br />

I thank the Royal Botanic Gardens Kew and the <strong>Center</strong><br />

<strong>for</strong> <strong>International</strong> <strong>Forestry</strong> <strong>Research</strong> <strong>for</strong> facilities and<br />

financial support<br />

References<br />

Berjak, P. and Pammenter, N.W. 1996. Recalcitrant<br />

(desiccation-sensitive) seeds. In: Olesen, K. (ed.)<br />

Innovations in tropical tree seed technology, 14-29.<br />

Danida Forest Seed Centre, Humlebaek.<br />

Chin, H.F., Hor, Y.L. and Mohd Lassim, M.B. 1984.<br />

Identification <strong>of</strong> recalcitrant seeds. Seed Science and<br />

Technology 12: 429-436.

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