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A review of dipterocarps - Center for International Forestry Research

A review of dipterocarps - Center for International Forestry Research

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Biogeography and Evolutionary Systematics <strong>of</strong> Dipterocarpaceae<br />

Figure 3. Paleogeographical reconstitution <strong>of</strong> southeast Asian Permo-Triassic in con<strong>for</strong>mity with paleobotanic<br />

data (from Vozenin-Serra and Salard-Cheboldaeff 1994).<br />

Be the origin in Europe or in Africa, both cases would<br />

have favoured dispersal and colonisation by the small,<br />

light, large-winged fruits over great distances and<br />

probably under drier conditions than those <strong>of</strong> the present<br />

rain <strong>for</strong>ests. This could explain why certain present taxa<br />

suggest an ancestral <strong>for</strong>m with these fruit characters and<br />

again brings <strong>for</strong>ward the hypothesis (Maury et al. 1975a,<br />

Maury 1978, 1979, Maury-Lechon 1979b) <strong>of</strong> an origin<br />

in open (perhaps semi-dry) environment. Forms with<br />

wingless fruits would have had the only possibility to<br />

concentrate their evolutionary potential into the<br />

protective structures around the seed (e.g., Vateria,<br />

Vateriopsis, Stemonoporus, certain Vatica and other<br />

9<br />

1. Sinkiang block (Tarim) + Sino-Korean block (North China).<br />

2. Yangtse block (North China).<br />

3. North Tibet block (North Xizang + North-West Xunnan + Shan Plateau).<br />

4. Khorat-Kontum block (East Thailand, most <strong>of</strong> Laos, Cambodia and Vietnam, South <strong>of</strong> Song Ba suture).<br />

5. South Tibet block (Lhassa plate + lands located South <strong>of</strong> Yalu Tsangpo).<br />

6. Kashmir.<br />

7. Iran and Afghanistan (Helmand block).<br />

8. Turkey.<br />

9. Spain.<br />

8<br />

6<br />

7<br />

5<br />

1<br />

3<br />

2<br />

4<br />

20<br />

species with large wingless fruits), these structures could<br />

have favoured water dispersal. In the taxa possessing<br />

winged fruits the evolutionary potential might have<br />

remained available <strong>for</strong> new opportunities. Upuna and<br />

Monotes kerstingii are perhaps parallel in this respect,<br />

as they suggest the hypothesis that they represent similar<br />

situations developed in or near Laurasia (Upuna) and in<br />

Gondwana (Monotes kerstingii) during long periods <strong>of</strong><br />

supposed separation.<br />

Past Continental Changes and Floral Evolution<br />

Past flora also underline a connection between large<br />

floristic-climatic changes and the main known collisions

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