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A review of dipterocarps - Center for International Forestry Research

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Biogeography and Evolutionary Systematics <strong>of</strong> Dipterocarpaceae<br />

Table 3. (continued) Affinities between Dipterocarpaceae sensu lato and other close angiosperm families.<br />

BOTANICAL CHARACTERS<br />

leaves opposite - - - +<br />

alternate + + + + + + + +<br />

leaf venation prominent pinnate + + + +<br />

vertically transcurrent + + + +<br />

entirely transcurrent and + + + + + +<br />

presence <strong>of</strong> columns <strong>of</strong> sclerenchyme -<br />

indistinct leaf venation + + ±<br />

dentate leaves ±<br />

paired basal leaf nerves - - +<br />

stipule ± ± ± - - + + + +<br />

hypodermis (+papillose lower epiderms) - +<br />

1-2 layered hypodermis ± + +<br />

hair (various within a section) stellate ± - -<br />

tufted ± - +<br />

glandules ± - -<br />

complex indumentum + + + +<br />

geniculate petiole + + + +<br />

indumentum + complex anatomy petiole (Malvales type) + + +<br />

rays uniseriate - + - -<br />

biseriate - +<br />

multiseriate + - - -<br />

mixed uni/multiseriate + +<br />

presence <strong>of</strong> resins + + + -<br />

intercellular resin canals + - - + -<br />

mucilage canals in cortex and cells in the epidermis + + + - +<br />

elongate medullary mucilage cells + + - + +<br />

arrangement <strong>of</strong> bast fibres * into outwardly tapering wedges + + + +<br />

pith and primary cortex with indumentum + + + +<br />

anomocytic stomata + + + +<br />

complex petiolar vascular supply ± - - -<br />

chromosomes n=7 + + (+) (+) (+) - -<br />

n=11 + - - - -<br />

±: present and other possibilities;<br />

+: present;<br />

-: absent;<br />

Dipterocarpoideae<br />

Dipterocarpaceae*<br />

Monotoideae<br />

sensu lato<br />

Pakaraimoideae<br />

Guttiferae<br />

Theaceae<br />

Tiliaceae<br />

Elaeocarpaceae<br />

Ochnaceae<br />

(-) or (+): exceptions;<br />

* : adapted from Ashton 1982, Maury-Lechon 1979, and other<br />

works (see in text: Classification).<br />

Sarcolaenaceae<br />

10

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