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On the Structure of the Excretory Organs of Amphioxus. Part 2.—The ...

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190 EDWIN S. GOODRICH.<br />

focussed to <strong>the</strong> lower surface; <strong>the</strong> nuclei <strong>of</strong> <strong>the</strong> wall are again<br />

visible. There is no opening.<br />

Innumerable figures could be given <strong>of</strong> series <strong>of</strong> sectious all<br />

telling <strong>the</strong> same story. But <strong>the</strong> critic will say : if <strong>the</strong> diverticula<br />

are really closed, sections taken at right angles through<br />

<strong>the</strong>ir tip should show <strong>the</strong> tubes cut across embedded in <strong>the</strong><br />

thickness <strong>of</strong> <strong>the</strong> wall. Such sections are not difficult to find,<br />

and I figure several on Plates 12 and 13.<br />

Figs. 10 and 12 represent two consecutive sections across<br />

<strong>the</strong> tip <strong>of</strong> a branch. In <strong>the</strong> fh'sb are seen <strong>the</strong> tubes entering<br />

<strong>the</strong> wall, while <strong>the</strong> next (fig. 12) strikes <strong>the</strong> lumen. A small<br />

part <strong>of</strong> this figure is shown slightly diagrammatised (fig. 11)<br />

on a larger scale. Again three consecutive sections are<br />

drawn in figs. 15, 16, and 17. Here two sections cut through<br />

<strong>the</strong> solid wall before <strong>the</strong> lumen is reached. Lastly, fig. 18<br />

represents a section through two adjacent processes, one <strong>of</strong><br />

which has been cut so as to expose <strong>the</strong> lumen, while <strong>the</strong> o<strong>the</strong>r<br />

shows very clearly <strong>the</strong> soleuocyte tubes piercing <strong>the</strong> wall aud<br />

embedded in its cytoplasm.<br />

The evidence <strong>of</strong> all <strong>the</strong>se sections is quite unequivocal; it<br />

would serve no good purpose to multiply instances; <strong>the</strong>re is<br />

no opening, <strong>the</strong> wall is continuous, and is traversed by <strong>the</strong><br />

tubes <strong>of</strong> <strong>the</strong> solenocytes.<br />

But <strong>the</strong>re is o<strong>the</strong>r evidence <strong>of</strong> a different nature leading to<br />

<strong>the</strong> same conclusion. I have observed in a living nepliridium<br />

<strong>the</strong> fluid inside <strong>the</strong> nephridhil canal so compressed, perhaps<br />

by <strong>the</strong> overlying cover-glass, that it dilated <strong>the</strong> tip <strong>of</strong> <strong>the</strong><br />

diverticulum so as to give rise to a bulgiug vesicle at its<br />

extremity. Now, such a swelling could obviously not be<br />

formed if <strong>the</strong> tip were open.<br />

We may now turn to injectious to corroborate our view. I<br />

have receutly injected <strong>the</strong> dorsal hyperbranchial coslom with<br />

Indian ink. The minute black particles were held in suspension<br />

in sea-water. Such a fluid, if introduced with a<br />

hypodermic syringe, can be made to fill <strong>the</strong> coslom. It is<br />

clear that if <strong>the</strong> nephridium communicated with <strong>the</strong> coslom<br />

<strong>the</strong> ink would penetrate into <strong>the</strong> canal; this would happen

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