PART 1

three collections of C. stemonea tested (New York, Minnesota, Sweden). It runs about the same as
baeomyesic acid but the spot characteristics are different. Three collections of C. brunneola (Florida, Maine &
Sweden) contain no detectable substances, four contain baeomycesic and squamatic acids (Florida, New
York, North Carolina & Sweden). Tibell reported baeomycesic acid in1980 but rejected it in 1987. It seems
probable that his unknowns B & C are the same as the single unknown spot that I get with my TLC system.
However, it is clear that more work needs to be done to confirm this. The baeomycesic acid strain is known
from a single collection at Gold Head Branch State Park (Clay County) where the unknown substance strain
also occurs. Most specimens checked had asci in chains. Collections from Duval and Hernando counties had
asci arising singly.
Chaenotheca floridana R. C. Harris, sp. nov
Ab aliis acidum virensicum continentia, ascosporis grosse ornamentis, 4-5 µm diametro et apothecis
pallidis brevibus et crassiusculis differt.
Type. FLORIDA. Putnam County: Ocala National Forest, old Johnson Field Campground along Oklawaha River
just W of U.S. Hwy. 19, swamp forest, 7 Dec 1988, Buck 16798 (NY, holotype; B, UPS, isotypes).
Thallus whitish, granular leprose, PD+ red-orange, virensic acid, unknown (tr.), protocetraric acid (tr.).
Phycobiont trebouxioid. Apothecia relatively short and stout, to 1 mm tall. Stalk to 0.3 mm across at base
tapering to ca. 0.1 mm, light orange brown (more or less concolorous with spore mass), appearing white
pruinose due to wefts of loose hyphae or occasionally covered with thallus particles. Capitulum obconical to
globose, to 0.5 mm in diameter. Asci not in chains. Ascospores globose or appearing slightly angular due to
coarse plate-like ornamentation, 4-5 µm in diameter, yellow brown.
I originally included this species in C. brunneola s. lat. until I studied its chemistry. A subsequent reevaluation
showed that it also differs in the pale, shorter apothecia and slightly larger, coarsely ornamented
ascospores.
Additional specimens. FLORIDA. Hillsborough County: Hillsborough River State Park, Rapids Trail
along river, 28°09'N, 82°14W, on trunk of Sabal, 3 Dec 1992, Buck 22641; Levy County: Cedar Key Scrub
State Preserve, along Co. Rd. 347 ca. 1.5 mi N of Fla. Hwy. 24, on Sabal leaf bases, 30 Nov 1992, Buck
22425, 22434, 22436; Sarasota County: Myakka River State Park, along Fla. Hwy. 72 ca. 0.7 mi SE of park
entrance, on trunk of Sabal, 5 Dec 1992, Buck 22717, 22731 (all NY).
Chaenotheca hispidula (Ach.) Zahlbr. s. lat.
The single specimen varies from the North American norm in having the asci in chains and in smaller
ascospores reaching only 4 µm which is below the minimum given by Tibell (1980). It is possible that the
deviation is clinal as the locality is considerably south of the range given by Tibell.
THELOMMA Massal.
Thelomma carolinianum (Tuck.) Tibell is endemic to the Coastal Plain. The other species of the
genus occur in western North America and Europe. See key to genera.
TYLOPHORON Nyl. ex Stizenb.
1. Thallus and ascomatal margin C+ red (lecanoric acid); ascomata
0.6-1.2 mm diam. (Tibell, 1982); ascospores 12-15 × 6.5-8 µm;
sporodochia white; conidiospores 1-2(-4)-celled, colorless,
rough-walled; throughout Florida.................................................................................. T. protrudens Nyl.
1. Thallus C-; ascomatal margin C+ red (lecanoric acid, 2'-O-methylperlatolic
acid); ascomata 0.4-0.8 mm diam. (Tibell, 1982); ascospores 10-14 × 5-6 µm; sporodochia
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