PART 1
PART 1
PART 1
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16. Thallus UV-................................................................................................................................................................................... 18<br />
18. Ascospores 14-19-celled, 55-80 µm long .............................................................................................................................. 19<br />
19. Stictic acid; exciple strongly developed below; ascospores ca. 14-celled,<br />
55 × 8-9 µm; Southwest Florida, Mar 1878, Smith & Austin 91.................................................................. Graphis sp. 91<br />
19. Norstictic acid; exciple barely closed below; ascospores 16-19-celled,<br />
70-80 × 10-11 µm; Louisiana, not yet known from Florida ..................................................[G. lumbricina Vainio s. lat.]<br />
18. Ascospores 8-12-celled, 20-45 µm long ................................................................................................................................ 20<br />
20. Margin carbonized to base of hymenium.......................................................................................................................... 21<br />
21. No substances; ascospores 10-12-celled, 35-45 × 7-9 µm;<br />
Duval and Lake counties....................................................................................................G. striatula (Ach.) Sprengel<br />
21. Stictic acid; ascospores 8-celled, 23-28 × 6-7 µm; Leon County .................................................... Graphis sp. 25050<br />
20. Margin carbonized above, orange below; ascospores 9-10-celled,<br />
25-27 × 6-7 µm often sterile; no substances; Duval and Marion counties ............................................G. subelegans Nyl.<br />
1. Exciple not carbonized; ascospores 2-4-celled .................................................................................................................................... 22<br />
.<br />
22. Ascospores 4-celled; no substances ............................................................................................................................................... 23<br />
23. Lirellae fissurine; exciple not evident.......................................................................................................................................... 24<br />
24. Margin not swollen; ascospores to 17 × 7 µm ....................................................................................................................... 25<br />
25. Thallus white, ecorticate; lirellae not gaping, minute, visible only as<br />
thin lines; ascospores 15-17 × 6-7 µm; Duval and Polk counties.................................................G. illiterata R. C. Harris<br />
25. Thallus greenish, with shiny prosoplectenchymatous cortex;<br />
lirellae gaping; ascospores 11-13 × 5-6 µm; Levy County.................................................... G. subnitidula Nyl. ex Tuck.<br />
24. Margin swollen; ascospores 17-20 × 8-10 µm; Duval, Lake and<br />
Marion counties ......................................................................................................... G. insidiosa (Knight & Mitten) Hook. f.<br />
23. Lirellae long, dichotomously branched; disk hidden to exposed;<br />
exciple orangish, well developed laterally or at base ................................................................................................................. 26<br />
26. Disk not exposed; exciple well developed laterally ............................................................................................................... 27<br />
27. Margin not striate; ascospores 15-17 × 6-8 µm;<br />
Duval and Liberty counties........................................................................................................................G. subparilis Nyl.<br />
27. Margin striate; ascospores 22-24 × 10-13 µm; East Florida, 1877, Smith..................................................G. rufula Mont.<br />
26. Disk exposed; "exciple" well developed below hymenium;<br />
ascospores 10-13 × 5 µm; Collier, Dade and Osceola counties .................................................................G. grammatis Fée<br />
22. Ascospores 2-celled or 8-9-celled ................................................................................................................................................... 28<br />
28. Ascospores 2-celled; norstictic acid; thallus whitish, shiny, UV-; lirellae crowded, initially<br />
with closed ± pruinose lips, later disk exposed, pale, pruinose; Duval County................................................ G. turbulenta Nyl.<br />
28. Ascospores 8-9-celled, 21-31 × 7.5-9 µm; thallus whitish, dull, UV+ yellow (lichexanthone);<br />
lirellae imnersed; disk pale red brown; margin white, patchily developed; Bay, Jefferson and<br />
Lafayette counties ......................................................................................................................................G. anguinoides ined.<br />
NOTES<br />
Graphis "anguinoides" ined.<br />
This species is superficially indistinguishable from Graphina anguina. I had actually placed a couple specimens in this<br />
species on the external characters of ascoma and UV+ thallus. When I checked the ascospores of some recent collections, I<br />
discovered a taxon with transversely septate, not muriform, ascospores. To my mind this is another nail in the coffin of the<br />
ascospore genera, Graphis vs. Graphina. It is inconceivable to me that genera defined on the single character of ascospore<br />
septation should have evolved parallel species with identical multicharacter complexes of thallus and ascoma.<br />
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