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Amphibalanus improvisus Print Date: 9/9/2012 Estuary P ECOSYSTEM Unconsolidated X Consolidated X Unvegetated X SAV MAR MAN D RI X SR CR O/M UV-CS UV-TF UV-SUB X TP RI-PH X X Coastal Fringe Epi-Surface PAR Mud Coastal Bay P Nearshore O Epipelagic Epi-Shallow Shelf DEPTH [Obs: 0 - 90m] [Pref: 0 - 6m] Benthic Depth Intertidal Subtidal P Bathyal P Sub-Shallow Sub-Deep P O Epi-Deep REGIME Oceanic Coastal Fringe Pelagic Depth Meso Bathyal Rivers O Lakes Abyssal UNCONSOLIDATED SUBSTRATE [Obs % Fines: 2.81 - 56.58%] X Sand Mix. Fines Gravel Cobble Mixed Sediments O Abyssal Terrestrial Hadal Hadal SALINITY [Obs: 0 - 40psu] [Pref: 6 - 30psu] Fresh O Brackish P Marine O Hyper O Oligohaline O Mesohaline P Polyhaline P Beta Alpha Beta Alpha Beta Alpha Beta Alpha O O O O P P P P K Pelagic Organic CONSOLIDATED SUBSTRATE X R P HP Biogenic P Artificial Substrate P C O M W CA K RA MAN DW R P H + B Oth P P P P P P O SynH X SA PP EARLY DEVELOPMENT V OVI OVO X PL 1265 Supralittoral H TROPHIC LEVEL AND FEEDING P S DET DEC SF X REPRODUCTION Sexual X Asexual H X G/D SF X BF BUD PAR/AGA SeqH IF X FEE FCS P DD DF DF-SUR DF-SUB JUVENILE DEVELOPMENT/DISPERSAL LP X FR SD LP-B LP-P X HABITAT ASSOCIATION Pelagic Benthic X Epibiotic X NE SUB DEM SUR X UR B N SF IN EPP EPZ BP EPS EPU EPC X X X F X VP SP SP
Amphibalanus reticulatus Taxon: Barnacle Kingdom: Animalia Subkingdom: Eumetazoa Phylum: Arthropoda Subphylum: Crustacea Superclass: Class: Maxillopoda Subclass: Thecostraca Infraclass: Cirripedia Superorder: Thoracica Order: Sessilia Suborder: Balanomorpha Infraorder: Superfamily: Balanoidea Family: Balanidae Subfamily: Also Known As (Name - Type): Common Names: Balanus reticulates Misspelling Balanus reticulatus Synonym Type Locality: Ecoregion-level distribution Master Classifications Date 1st record: 1934 Loc 1st record: Japan Established: Yes NWP Taxonomic Author: (Utinomi, 1967) Hawaii 1929 Oahu, Hawaii Yes Species ID: 15439 Print Date: 9/9/2012 10:47:37 PM NEP 1984 Mazatlan, Mexico Yes VECTORS SH X MS AF ID RE AP REC SF HR O BW SB HF S/R AE AA IR A P X X AO PO Comments: Amphibalanus reticulatus was considered native in Japan (Iwasaki, 2006); however Doi et al. (2011) list it as cryptogenic. We classify it as a conflict in the ecoregions boarding mainland Japan and native in the Western and Central Indo-Pacific with the exception of Australia where it is introduced (Jones, 2004) or unclassified. 1266
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DISCLAIMER: The information in this
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TABLE OF CONTENTS Disclaimer ......
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Undaria pinnatifida ...............
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Diadumene leucolena ...............
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Hyboscolex longiseta ..............
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Watersipora subtorquata ...........
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Littoridinops monroensis ..........
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Halophiloscia couchii .............
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Amphibalanus amphitrite ...........
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Botrylloides diegensis ............
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Salmo trutta ......................
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Figure 45: Mollusca-Gastropoda: Num
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INTRODUCTION Overview Marine and es
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APPROACH AND DEFINITIONS Marine Bio
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Unknown: A species was classified a
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new species. In these cases, the pa
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accumulate in rocky intertidal tide
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the second most invaded. The Northe
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history of the species but not whet
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MAPS OF MEOW REGIONS Figure 1: Map
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Figure 3: Maps in geographic projec
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Figure 5: Map of the Northwest Paci
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Figure 7: Map of the Eastern Indo-P
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Figure 9: Map of the Tropical Easte
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Figure 11: Map of the Magellanic re
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Figure 13: Map of the West Tropical
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Figure 15: Map of the Northeast Atl
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Figure 17: Map of the Ponto-Caspian
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Figure 19: Map of the Southern Afri
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Figure 21: Map of the Southern Aust
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Figure 23: Alternate map in polar s
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250 200 150 100 50 0 N = 747 10 55
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25 20 15 10 2 6 Figure 30: Phylum B
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6 5 4 3 2 1 0 0 0 Figure 38: Cnidar
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140 120 100 Figure 46: Phylum Arthr
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Figure 54: Arthropoda-Insecta: Numb
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DECIPHERING SPECIES INFORMATION: KE
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TABLES Table Key to Species Profile
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Attribute / Level in Species Profil
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Level 1 Level 2 Level 3 Definition
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Ecosystems Table 4: Definitions for
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Major Habitat Type: Level 1 Ecosyst
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Depth Table 5: Definitions for the
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Level 1 Level 2 Level 3 Definition
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Salinity Table 7: Definitions for t
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Level 1 Level 2 Level 3 Level 4 Exp
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Level 1 Level 2 Level 3 Level 4 Exp
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Juvenile Development/Dispersal Leve
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Level 1 Level 2 Level 3 Level 4 Def
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Level 1 Level 2 Level 3 Level 4 Def
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Claviceps purpurea var. spartinae P
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Cytospora rhizophorae Print Date: 9
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Etheirophora blepharospora Print Da
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Bonamia ostreae Print Date: 9/9/201
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Boveria teredinidi Print Date: 9/9/
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Cothurnia limnoriae Print Date: 9/9
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Eufolliculina lignicola Print Date:
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Lobochona prorates Print Date: 9/9/
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Mirofolliculina limnoriae Print Dat
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Haynesina germanica Print Date: 9/9
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Attheya armata Print Date: 9/9/2012
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Pseudo-nitzschia americana Print Da
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Saccharina japonica Print Date: 9/9
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Striaria attenuata Print Date: 9/9/
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Undaria pinnatifida Print Date: 9/9
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Dasya sessilis Print Date: 9/9/2012
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Polysiphonia denudata Print Date: 9
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Bryopsis sp. (Cohen and Carlton, 19
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Udotea argentea Print Date: 9/9/201
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Ulva californica Print Date: 9/9/20
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Ulva expansa Print Date: 9/9/2012 E
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Ulva lactuca Print Date: 9/9/2012 E
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Ulva pertusa Print Date: 9/9/2012 E
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Ulva rigida Print Date: 9/9/2012 Es
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Ammophila arenaria Print Date: 9/9/
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Halophila decipiens Print Date: 9/9
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Limosella australis Print Date: 9/9
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Rhizophora mangle Print Date: 9/9/2
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Monanchora sp. (Carlton and Eldredg
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Mycale cecilia Print Date: 9/9/2012
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Mycale grandis Print Date: 9/9/2012
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Mycale parishi Print Date: 9/9/2012
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Aiptasia cf. insignis Print Date: 9
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Aiptasiomorpha minima Print Date: 9
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Bunodeopsis sp. A (Engle and Richar
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Carijoa riisei Print Date: 9/9/2012
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Culicia rachelfitzhardingeae Print
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Diadumene lineata Print Date: 9/9/2
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Synandwakia hozawai Print Date: 9/9
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Copula sivickisi Print Date: 9/9/20
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Amphinema sp. (Rees, 2000) Print Da
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Bimeria vestita Print Date: 9/9/201
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Blackfordia virginica Print Date: 9
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Bougainvillia muscus Print Date: 9/
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Cladonema pacificum Print Date: 9/9
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Cladonema radiatum Print Date: 9/9/
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Clava multicornis Print Date: 9/9/2
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Clytia hemisphaerica Print Date: 9/
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Cordylophora caspia Print Date: 9/9
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Corymorpha sp. (Cohen and Carlton,
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Garveia franciscana Print Date: 9/9
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Garveia sp. (Carlton and Eldredge,
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Gonothyraea loveni Print Date: 9/9/
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Laomedea flexuosa Print Date: 9/9/2
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Maeotias marginata Print Date: 9/9/
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Moerisia sp. (Rees and Gershwin, 20
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Obelia bidentata Print Date: 9/9/20
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Obelia dichotoma Cmplx Print Date:
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Pennaria disticha Print Date: 9/9/2
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Pinauay crocea Print Date: 9/9/2012
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Podocorynoides minima Print Date: 9
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Thuiaria thuiarioides Print Date: 9
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Turritopsis nutricula Cmplx Print D
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Anomalorhiza shawi Print Date: 9/9/
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Aurelia sp. (Carlton and Eldredge,
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Aurelia sp. 1 (Dawson et al., 2005)
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Cassiopea andromeda Print Date: 9/9
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Cassiopea sp. 3 (Holland et al., 20
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Vallicula multiformis Print Date: 9
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Cercaria batillariae Print Date: 9/
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Khawia iowensis Print Date: 9/9/201
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Taenioplana teredini Print Date: 9/
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Branchiomma bairdi Print Date: 9/9/
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Clymenella torquata Print Date: 9/9
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Crucigera websteri Print Date: 9/9/
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Demonax sp. A (Cohen et al., 2002)
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Dipolydora normalis Print Date: 9/9
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Eumida sanguinea Print Date: 9/9/20
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Eusyllis japonica Print Date: 9/9/2
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Exogone longicornis Print Date: 9/9
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Myrianida pachycera Print Date: 9/9
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Nicolea sp. A (Cohen et al., 2002)
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Spio blakei Print Date: 9/9/2012 Es
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Amathia convoluta Print Date: 9/9/2
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Bowerbankia imbricata Print Date: 9
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Bugula californica Print Date: 9/9/
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Bugula dentata Print Date: 9/9/2012
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Bugula flabellata Print Date: 9/9/2
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Bugula fulva Print Date: 9/9/2012 E
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Bugula minima Print Date: 9/9/2012
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Bugula neritina Cmplx Print Date: 9
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Bugula sp. 1 (Cohen et al., 1998) P
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Bugula sp. 2 (Cohen et al., 1998) P
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Escharoides excavata Print Date: 9/
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Nolella stipata Print Date: 9/9/201
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Synnotum aegyptiacum Print Date: 9/
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Triticella sp. B (Hewitt, 1993) Pri
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Watersipora arcuata Print Date: 9/9
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Hiatella arctica Print Date: 9/9/20
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Limnoperna fortunei Print Date: 9/9
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Lioconcha fastigiata Print Date: 9/
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Macoma petalum Print Date: 9/9/2012
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Martesia striata Print Date: 9/9/20
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Mya arenaria Print Date: 9/29/2012
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Ostrea edulis Print Date: 9/9/2012
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Ostrea lurida Print Date: 9/9/2012
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Panopea generosa Print Date: 9/9/20
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Patinopecten yessoensis Print Date:
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Perna viridis Print Date: 9/9/2012
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Teredo bartschi Print Date: 9/9/201
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Theora lubrica Print Date: 9/9/2012
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Alderia modesta Print Date: 9/9/201
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Babakina festiva Print Date: 9/9/20
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Batillaria attramentaria Print Date
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Boonea cincta Print Date: 9/9/2012
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Bostrycapulus calyptraeiformis Prin
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Caloria indica Print Date: 9/9/2012
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Catriona rickettsi Print Date: 9/9/
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Cecina manchurica Print Date: 9/9/2
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Chrysallida trachis Print Date: 9/9
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Crepidula convexa Print Date: 9/9/2
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Crepidula fornicata Print Date: 9/9
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Crepidula onyx Print Date: 9/9/2012
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Crucibulum spinosum Print Date: 9/9
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Cuthona columbiana Print Date: 9/9/
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Cuthona perca Print Date: 9/9/2012
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Diodora rueppellii Print Date: 9/9/
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Folinella navisa Print Date: 9/9/20
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Haliotis fulgens Print Date: 9/9/20
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Haliotis iris Print Date: 9/9/2012
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Haliotis kamtschatkana Print Date:
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Haliotis laevigata Print Date: 9/9/
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Haliotis rufescens Print Date: 9/9/
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Haminoea japonica Print Date: 9/9/2
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Ilyanassa obsoleta Print Date: 9/9/
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Iolaea eucosmia Print Date: 9/9/201
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Laguncula pulchella Print Date: 9/9
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Littoridinops monroensis Print Date
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Littorina littorea Print Date: 9/9/
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Littorina saxatilis Print Date: 9/9
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Myosotella myosotis Print Date: 9/9
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Nassarius fraterculus Print Date: 9
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Nassarius sinarus Print Date: 9/9/2
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Ocenebra inornata Print Date: 9/9/2
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Odetta bisuturalis Print Date: 9/9/
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Okenia eolida Print Date: 9/9/2012
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Okenia pellucida Print Date: 9/9/20
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Peristichia pedroana Print Date: 9/
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Philine auriformis Print Date: 9/9/
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Philine orientalis Print Date: 9/29
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Phyllodesmium poindimiei Print Date
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Pyrgophorus coronatus Print Date: 9
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Sakuraeolis enosimensis Print Date:
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Stenothyra sp. (Tamaki et al., 2002
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Tarebia granifera Print Date: 9/9/2
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Tenellia adspersa Print Date: 9/9/2
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Thylaeodus sp. (Carlton and Eldredg
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Umbonium thomasi Print Date: 9/9/20
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Urosalpinx cinerea Print Date: 9/9/
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Vermetus alii Print Date: 9/9/2012
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Nebalia sp. (Cohen and Carlton, 199
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Ampelisca abdita Print Date: 9/9/20
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Ampithoe lacertosa Print Date: 9/9/
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Ampithoe longimana Print Date: 9/9/
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Ampithoe sp. (Cohen and Carlton, 19
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Ampithoe valida Print Date: 9/9/201
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Aoroides secundus Print Date: 9/9/2
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Caprella danilevskii Print Date: 9/
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Caprella drepanochir Print Date: 9/
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Caprella equilibra Print Date: 9/9/
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Caprella mutica Print Date: 9/9/201
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Caprella penantis Print Date: 9/9/2
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Caprella scaura Print Date: 9/9/201
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Caprella simia Print Date: 9/9/2012
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Centromedon pumilus Print Date: 9/9
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Chelura terebrans Print Date: 9/9/2
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Corophium alienense Print Date: 9/9
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Corophium heteroceratum Print Date:
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Elasmopus rapax Print Date: 9/9/201
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Eochelidium sp. A (Chapman, 2007) P
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Ericthonius brasiliensis Print Date
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Gammarus daiberi Print Date: 9/9/20
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Gammarus mucronatus Print Date: 9/9
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Grandidierella bispinosa Print Date
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Incisocalliope derzhavini Print Dat
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Jassa falcata Print Date: 9/9/2012
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Jassa marmorata Print Date: 9/9/201
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Jassa slatteryi Print Date: 9/9/201
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Laticorophium baconi Print Date: 9/
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Leucothoe micronesiae Print Date: 9
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Leucothoe spinicarpa Cmplx Print Da
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Liljeborgia sp. (Cohen et al., 2002
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Melita nitida Print Date: 9/9/2012
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Melita rylovae Print Date: 9/9/2012
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Metopella sp. (Cohen et al., 2002)
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Microdeutopus gryllotalpa Print Dat
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Microjassa sp. (deRivera et al., 20
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Monocorophium uenoi Print Date: 9/9
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Paracaprella pusilla Print Date: 9/
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Paracorophium sp. (Chapman, 2007) P
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Paradexamine sp. (Chapman, 2007) Pr
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Paraleucothoe sp. (Carlton and Eldr
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Stenothoe gallensis Print Date: 9/9
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Stenothoe valida Print Date: 9/9/20
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Transorchestia enigmatica Print Dat
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Tropichelura insulae Print Date: 9/
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Nannastacus sp. (Carlton and Eldred
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Nippoleucon hinumensis Print Date:
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Scherocumella sp. (Carlton and Eldr
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Alloniscus oahuensis Print Date: 9/
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Armadilloniscus ellipticus Print Da
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Boreosignum wilsoni Print Date: 9/9
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Buchnerillo sp. (Carlton and Eldred
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Caecijaera horvathi Print Date: 9/9
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Dynoides dentisinus Print Date: 9/9
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Eurylana arcuata Print Date: 9/9/20
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Exosphaeroma sp. (Carlton and Eldre
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Gnorimosphaeroma rayi Print Date: 9
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Halophiloscia couchii Print Date: 9
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Iais californica Print Date: 9/9/20
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Ligia exotica Print Date: 9/9/2012
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Limnoria quadripunctata Print Date:
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Limnoria tripunctata Print Date: 9/
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Littorophiloscia culebrae Print Dat
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Mesanthura sp. (Carlton and Eldredg
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Munna sp. A (Cohen and Carlton, 199
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Niambia capensis Print Date: 9/9/20
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Olibrinus truncatus Print Date: 9/9
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Orthione griffenis Print Date: 9/9/
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Paracerceis sculpta Print Date: 9/9
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Paradella dianae Print Date: 9/9/20
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Paralimnoria andrewsi Print Date: 9
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Paranthura japonica Print Date: 9/9
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Pistorius bidens Print Date: 9/9/20
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Platyarthrus aiasensis Print Date:
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Porcellio dilatatus Print Date: 9/9
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Porcellio laevis Print Date: 9/9/20
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Porcellio lamellatus Print Date: 9/
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Porcellio scaber Print Date: 9/9/20
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Pseudosphaeroma sp. (Bruce and Wetz
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Sphaeroma quoianum Print Date: 9/9/
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Sphaeroma walkeri Print Date: 9/9/2
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Synidotea laevidorsalis Print Date:
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Deltamysis holmquistae Print Date:
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Holmesimysis costata Print Date: 9/
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Hyperacanthomysis longirostris Prin
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Neomysis japonica Print Date: 9/9/2
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Orientomysis aspera Print Date: 9/9
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Orientomysis hwanhaiensis Print Dat
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Apseudes sp. (Carlton and Eldredge,
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Parapseudes pedispinis Print Date:
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Sinelobus stanfordi Cmplx Print Dat
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Acantholobulus pacificus Print Date
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Callinectes sapidus Print Date: 9/9
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Carcinus aestuarii Print Date: 9/9/
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Carcinus maenas Print Date: 10/2/20
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Deiratonotus cristatum Print Date:
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Eriocheir sinensis Print Date: 9/9/
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Exopalaemon carinicauda Print Date:
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Fenneropenaeus chinensis Print Date
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Fenneropenaeus merguiensis Print Da
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Homarus americanus Print Date: 9/9/
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Homarus gammarus Print Date: 9/9/20
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Hyastenus spinosus Print Date: 9/9/
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Litopenaeus vannamei Print Date: 9/
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Macrobrachium lar Print Date: 9/9/2
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Macrobrachium rosenbergii Print Dat
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Marsupenaeus japonicus Print Date:
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Metacarcinus magister Print Date: 9
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Metadromia wilsoni Print Date: 9/9/
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Nanosesarma minutum Print Date: 9/9
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Pachygrapsus fakaravensis Print Dat
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Palaemon macrodactylus Print Date:
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Panopeus lacustris Print Date: 9/9/
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Pilumnus oahuensis Print Date: 9/9/
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Pyromaia tuberculata Print Date: 9/
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Scylla serrata Print Date: 9/9/2012
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Page 1260 and 1261: Oithona davisae Print Date: 9/9/201
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Page 1276 and 1277: Sinocalanus doerrii Print Date: 9/9
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Page 1306 and 1307: Balanus perforatus Print Date: 9/9/
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Page 1318 and 1319: Aspidoconcha limnoriae Print Date:
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Page 1328 and 1329: Ammothella pacifica Print Date: 9/9
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Endeis biseriata Print Date: 9/9/20
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Endeis nodosa Print Date: 9/9/2012
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Endeis procera Print Date: 9/9/2012
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Pigrogromitus timsanus Print Date:
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Tanystylum rehderi Print Date: 9/9/
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Anisolabis maritima Print Date: 9/9
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Atrichopogon jacobsoni Print Date:
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Atrichopogon sp. (Carlton & Eldredg
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Brachydeutera ibari Print Date: 9/9
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Canaceoides angulatus Print Date: 9
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Cercyon fimbriatum Print Date: 9/9/
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Ceropsilopa coquilletti Print Date:
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Clasiopella uncinata Print Date: 9/
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Cricotopus bicinctus Print Date: 9/
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Crocothemis servilia Print Date: 9/
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Cyclodinus mundulus Print Date: 9/9
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Dolichopus exsul Print Date: 9/9/20
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Donaceus nigronotatus Print Date: 9
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Enallagma civile Print Date: 9/9/20
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Enochrus sayi Print Date: 9/9/2012
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Ephydra gracilis Print Date: 9/9/20
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Ephydra millbrae Print Date: 9/9/20
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Goeldichironomus holoprasinus Print
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Hecamede granifera Print Date: 9/9/
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Hostis guamensis Print Date: 9/9/20
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Ischnura ramburii Print Date: 9/9/2
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Kleidotoma bryani Print Date: 9/9/2
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Medetera grisescens Print Date: 9/9
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Mesovelia amoena Print Date: 9/9/20
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Mesovelia mulsanti Print Date: 9/9/
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Micracanthia humilis Print Date: 9/
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Mosillus tibialis Print Date: 9/9/2
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Orasiopa mera Print Date: 9/9/2012
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Parathroscinus murphyi Print Date:
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Paratissa pollinosa Print Date: 9/9
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Placopsidella marquesana Print Date
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Procanace williamsi Print Date: 9/9
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Prokelisia marginata Print Date: 9/
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Pselactus spadix Print Date: 9/9/20
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Psilopa girschneri Print Date: 9/9/
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Psychoda salicornia Print Date: 9/9
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Scatella stagnalis Print Date: 9/9/
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Syntormon flexible Print Date: 9/9/
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Telmatogeton japonicus Print Date:
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Tethina willistoni Print Date: 9/9/
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Thambemyia borealis Print Date: 9/9
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Tramea lacerata Print Date: 9/9/201
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Trichocorixa reticulata Print Date:
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Trigonotylus uhleri Print Date: 9/9
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Tropisternus salsamentus Print Date
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Ophiactis savignyi Print Date: 9/9/
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Strongylocentrotus intermedius Prin
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Ascidia archaia Print Date: 9/9/201
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Ascidia sp. (Lambert and Lambert, 1
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Ascidia species A (Carlton and Eldr
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Ascidia species B (Carlton and Eldr
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Ascidia sydneiensis Print Date: 9/9
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Styela clava Print Date: 9/9/2012 E
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Symplegma reptans Print Date: 9/9/2
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Cyprinus carpio Print Date: 9/9/201
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Ictalurus furcatus Print Date: 9/9/
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Lateolabrax sp. (Yokogawa and Seki,
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Lates calcarifer Print Date: 9/9/20
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Limia vittata Print Date: 9/9/2012
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Lucania parva Print Date: 9/9/2012
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Perca flavescens Print Date: 9/9/20
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Poecilia mexicana Print Date: 9/9/2
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Poecilia reticulata Print Date: 9/9
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Psetta maxima Print Date: 9/9/2012
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Upeneus vittatus Print Date: 9/9/20
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Valamugil engeli Print Date: 9/9/20
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Ahyong, S.T. 2001. Revision of the
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Aquenal Pty. Ltd. 2002. Final repor
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Ayres, P. 1991. Introduced Pacific
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Beasley, C.R., Fernandes, C.M., Gom
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Blake, J.A., Hecker, B., Grassle, J
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Brasher, A.M.D., Luton, C.D., Goodb
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Busby, M.S., Barnhart, R.A., and Pe
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Carlton, J.T. 1987. Patterns of tra
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Castanos, C., Pascual, M., and Cama
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Chen, R., Watanabe, S., and Yokota,
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Wastewater Department, Environmenta
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Cragg, S.M., Jumel M.-C., Al-Horani
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Daly, H.V. 2007a. Orders of interti
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Farrapeira, C.M.R. 2010b. Shallow w
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Godwin, L.S. 2003. Hull fouling of
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Ho, J. 2005. Symbiotic copepods as
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Lewis, J.E. and Norris, J.N. 1987.
Page 1748 and 1749:
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Marquet, G., Taiki, N., Chadderton,
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Morton, A. and Volpe, J. 2003. A de
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Rizzo, A.E. and Amaral, A.C.Z. 2001
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Ruiz, G.M., Fofonoff, P.W., Carlton
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SCAMIT (Southern California Associa
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Senanan, W., Tangkrock-Olan, N., Pa
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Silva, P.C., Basson, P.W., and Moe,
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Toda, H. and Yamamoto, T. 1991. Dis
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