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McGUIRE-SYSTEMATICS OF CROTAPHYTID LIZARDS<br />
tographs were provided in Avila (1 995) and Sprack-<br />
land (1993).<br />
Crof aphytus grismeri McGuire<br />
(Fig. 32B)<br />
Croraphytus grisnieri McGuire, 1994:439; fig. I. Type locality:<br />
"Cation David, a low pass that separates the contiguous Sierra<br />
de Los Cucapas and Sierra El Mayor, approximately 2 km W<br />
Mex. Hwy. 5 on the dirt road to the sulfur mine (turnoff at km<br />
49 S. Mexicali), Baja California, Mexico" (holotype: CES 067-<br />
629).<br />
Etymology.-Named in honor of L. Lee Grismer, noted au-<br />
thority on the herpetology of Baja California.<br />
Diagnosis. - Crotaphytus grismeri differs from all<br />
other Crotaphytus in the presence of a dull orange<br />
colored tail and hind limbs in subadult females,<br />
green pigmentation within the pale gray or white bar<br />
that separates the anterior and posterior black col-<br />
lars, a well-defined pale tan dorsal caudal stripe in<br />
juveniles of both sexes, a hindlimb pattern wherein<br />
the region between the middle of the thigh and its<br />
distal extremity is yellow and unmarked except for<br />
scattered minute brown spots, and in its small adult<br />
size (maximum male SVL = 99 mm, n = 7; X =<br />
93.3). The presence in subadult females (n = 6, in-<br />
cluding photographs of living individuals) of three<br />
large, lateral black spots with bold white borders<br />
may represent another diagnostic feature. Crota-<br />
phytus grismeri is further distinguished from C. re-<br />
riculatus, C. antiquus, and C. collaris by the pres-<br />
ence, in adult males, of large black or dark brown<br />
inguinal patches, a strongly laterally compressed tail,<br />
and a bold white dorsal caudal stripe. It differs from<br />
these species and from C. dickersonaein that it lacks<br />
(in both sexes) black pigmentation of the oral mu-<br />
cosa and in the dark brown dorsal ground color of<br />
adult males. It differs from the remaining Crota-<br />
phytus (C. bicinctores, C. insularis, and C. vestigium)<br />
in that the dorsal surface of the forelimb is yellow<br />
and almost without pattern, except for a small patch<br />
of minute brown spots near the forelimb insertion.<br />
It differs further from C. insularis and C. vestigium<br />
in that the posterior collar is only narrowly incom-<br />
plete middorsally rather than broadly incomplete<br />
and in having a dorsal pattern of subequal white<br />
spots without transversely oriented white bars.<br />
Variation (n = 10). - Rostral approximately four<br />
times wider than high, usually rectangular in shape.<br />
Rostral bordered by three to four postrostrals. Re-<br />
maining snout scales irregularly arranged, an en-<br />
larged middorsal series may be present. Nasals sep-<br />
arated by five to six internasals. Frontonasals oc-<br />
casionally enlarged. Canthals three; five to seven<br />
scales separate canthals of left and right sides. Su-<br />
praorbital semicircles present with ten to 15 scales<br />
per semicircle, median scales sometimes fuse to form<br />
an azygous frontal. Supraoculars flat or convex,<br />
smooth, becoming progressively larger medially such<br />
that medial scales are two to four times larger than<br />
lateral ones. Circumorbitals present, not well dif-<br />
ferentiated from supraoculars. Superciliaries nine to<br />
13, elongate medial scale occasionally present. Pal-<br />
pebrals ovoid, slightly convex, interspersed with nu-<br />
merous interstitial granules. Preoculars, suboculars,<br />
and postoculars form an arc of seven to 13 rectan-<br />
gular scales, the second, third, or fourth scale only<br />
rarely elongate. Supralabials 14 to 17, usually slight-<br />
ly longer than high except anteriormost scale, which<br />
is square or pentagonal. Lorilabials in two or three<br />
rows, ovoid to rectangular, juxtaposed, separating<br />
supralabials from suboculars and nasals. Aperture<br />
of external auditory meatus rectangular or ovoid,<br />
often constricted at or above the midpoint, approx-<br />
imately four times higher than wide, with small,<br />
strongly convex, somewhat conical auricular scales<br />
lining anterior margin. Mental pentagonal, one to<br />
1.5 times wider than high, bordered laterally by an-<br />
terior infralabials and posteriorly by a pair of large<br />
postmentals. Postmentals may or may not be sep-<br />
arated from infralabials by a pair of sublabials.<br />
Chinshields weakly differentiated or undifferentiat-<br />
ed. Infralabials 13 to 18, square or wider than high,<br />
inferior border convex. Gulars granular, strongly<br />
convex and beadlike, each scale separated from ad-<br />
jacent scales by numerous asymmetrically arranged<br />
interstitial granules.<br />
Dorsal scales in approximately 164 to 190 rows<br />
midway between forelimb and hindlimb insertions.<br />
Tail long, cylindrical to oval in females and juve-<br />
niles over entire length, anterior one-half strongly<br />
compressed laterally in adult males. Paired, median<br />
row of subcaudals larger than adjacent subcaudals<br />
and lateral caudals. Enlarged postanal scales present<br />
in males.<br />
Deep postfemoral dermal mite pocket present at<br />
hindlimb insertion. Femoral pores 19 to 23, femoral<br />
pores do not extend beyond angle of knee, separated<br />
medially by 20 to 25 granular scales. Subdigital la-<br />
mellae on fourth toe 18 to 20.<br />
Coloration in LiJe. -Dorsal body coloration in<br />
adult males is brown, without pale orange or peach<br />
colored body bands. The white component of the<br />
dorsal pattern is composed of white spots and oc-<br />
casional dashes on the body, as well as the proximal