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.................................................................................. .............................................................................. ......................................................................... ..................................................................... ....................................................................... Abstract Introduction Historical Review Materials and Methods Frequency Coding Allozyme Data Set ....................................................................... Ingroup Monophyly ...................................................................... Choice of Terminal Taxa ................................................................. Outgroup Taxa .......................................................................... Morphology and Character Descriptions ........................... . ....................... Skull Roof .............................................................................. Palate .................................................................................. Braincase ............................................................................... Mandible ............................................................................... Miscellaneous Features of the Head Skeleton ................................ . ............ Axial Skeleton .................... . ................................................... ...................... .......... .................................... Pectoral Girdle . . Pelvic Girdle ............................................................................ Limbs ................................... . ............................................ Squamation ............................................................................. Pockets and Folds ....................................................................... Additional Morphological Characters ............................. . ...................... Coloration .............................................................................. Behavioral Characters .................................................................... ............................................................................. ................................................................................... ................................................................................ Character List Results Discussion Comparison with Previous Hypotheses ................................ . ................. Character Evolution ..................................................................... Taxonomic Accounts ...................................................................... Crotaphytidae ........................................................................... Crotaphytus ........................................................................... C . antiquus ......................................................................... C . bicinctores ........................................................................ C . collaris ........................................................................... C . dickersonae ...................................................................... C . grismeri .......................................................................... C . insularis ......................................................................... C . nebrius .......................................................................... C . oligocenicust ..................................................................... C . reticulatus ........................................................................ C . vestigium ......................................................................... Gambelia ............................................................................. G.copei ............................................................................ G . coronat .......................................................................... G . silus ............................................................................. G . wislizenii ......................................................................... Key to the Species of Crotaphytus and Gambelia .............................................. Acknowledgments ......................................................................... Literature Cited ........................................................................... ... 111
..................................................... .................................................................. Appendices ...................... . 120 1 . Specimens Examined 120 2 . Data Matrix ........................ . ............................................. 126 3 . Outgroup Data Matrix ......................... . .. . ........................... 128 4 . Step Matrices for Manhattan Distance Frequency Approach ........................ . .... 132 5 . Character Transformations for Each Stem of the Single Most Parsimonious Tree 134 ............. 6 . List of Character State Changes by Character ............................................. 139 7 . Scleral Ossicle Data ................................................................... 143
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C. vestigiurn by the absence in adu
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