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The Geography of Phytochemical Races

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184 4 Intercontinental Disjunctions<br />

Fig. 4.7 Compounds 316–323, fl avonoids from some Restionaceae<br />

involving a much broader sampling has been described (Harborne, 1979). Fourteen<br />

Australasian species representing six genera and 33 species representing 10 African<br />

genera were subjected to chromatographic separation, and identifi cation <strong>of</strong> their fl avonoid<br />

glycosides. Flavonols identifi ed included quercetin, myricetin, the myricetin<br />

methyl ethers larycitrin [317] and syringetin [318], gossypetin [319], and gossypetin<br />

7-methyl ether [320]. Flavones identifi ed were apigenin, luteolin, chrysoeriol, and<br />

hypolaetin. C-Glycosylfl avones and proanthocyanidins were identifi ed from several<br />

species and anthocyanins from a few.<br />

<strong>The</strong> distribution <strong>of</strong> these compounds proved to be <strong>of</strong> considerable taxonomic<br />

value. <strong>The</strong> most interesting observation centered on the distribution <strong>of</strong> fl avonoids<br />

in members <strong>of</strong> the three disjunct genera. <strong>The</strong> pigment pr<strong>of</strong>i les <strong>of</strong> Australasian and<br />

African members <strong>of</strong> all three correlated with their geographic origins, and not with<br />

other members <strong>of</strong> their respective genera. A solution to this problem was soon<br />

to appear in the form <strong>of</strong> a total revision <strong>of</strong> the family (e.g., Briggs and Johnson,<br />

1998a, b). As mentioned by Williams et al. (1998), <strong>of</strong> the 147 species examined, 57<br />

are newly recognized, while 19 <strong>of</strong> 34 genera are new or involve resurrected names

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