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Middle Miocene palynoflora of the Legnica lignite deposit complex ...

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Cookson, Phragmothyrites Edwards, Plochmopeltinites<br />

Cookson, and Trichothyrites<br />

Rosendahl emend. Smith = Notothyrites Cookson)<br />

were found.<br />

The Microthyriaceae fungi are usually<br />

ectoparasites extended in tropical and subtropical<br />

regions. Their presence is an important<br />

index <strong>of</strong> high total annual rainfall – above<br />

1000 mm. They usually live on leaves <strong>of</strong> seed<br />

plants (Betulaceae, conifers and o<strong>the</strong>rs) and<br />

ferns – Pteridium, Aspidum and o<strong>the</strong>rs (Neuy-<br />

Stolz 1958, Elsik 1978). According to Elsik (op.<br />

cit.) <strong>the</strong>y occur in <strong>deposit</strong>s since <strong>the</strong> Lower Cretaceous,<br />

but <strong>the</strong> most numerous are Eocene<br />

and <strong>Miocene</strong> localities.<br />

ABRIDGED DESCRIPTION<br />

OF POLLEN DIAGRAMS<br />

The results <strong>of</strong> palynological investigations<br />

are presented in three pollen diagrams (Figs<br />

3–5) constructed using <strong>the</strong> POLPAL computer<br />

program (Nalepka & Walanus 2003). The percentage<br />

values are based on <strong>the</strong> total sum <strong>of</strong><br />

pollen grains and spores. Most taxa have <strong>the</strong>ir<br />

own columns. However, some columns present<br />

a sum <strong>of</strong> a few taxa (two or more genera, one<br />

family or two families). The sporomorphs are<br />

arranged in <strong>the</strong> diagrams according to plant<br />

communities from mesophytic to wet. Herbs,<br />

pteridophytes and mosses are presented separately<br />

at <strong>the</strong> end <strong>of</strong> <strong>the</strong> histograms.<br />

The pollen diagrams have been divided into<br />

four sections (L1–L4), containing samples with<br />

similar frequency <strong>of</strong> <strong>the</strong>rmophilous taxa.<br />

LUSATIAN SEAM<br />

This part (section L1) contains samples<br />

from <strong>the</strong> pr<strong>of</strong>i le <strong>Legnica</strong> 33/56 from <strong>the</strong> depth<br />

112.0–100.5 m (Nos 46–24) as well as <strong>Legnica</strong><br />

41/52 – depth 125.5–114.4 m (Nos. 43–20),<br />

taken mainly from brown coal. The bottom<br />

samples <strong>of</strong> both pr<strong>of</strong>i les, taken from light-grey<br />

sandy clay, could represent <strong>the</strong> Silesian-Lusatian<br />

series. However, ra<strong>the</strong>r bad preservation<br />

<strong>of</strong> sporomorphs makes impossible detailed<br />

studies and distinguishing a separate section<br />

for <strong>the</strong> Silesian-Lusatian series.<br />

Although <strong>the</strong> arctotertiary element prevails,<br />

<strong>the</strong> palaeotropical taxa play an important<br />

role (Tab. 1), and are represented mainly<br />

by Tricolporopollenites pseudocingulum (up to<br />

65<br />

42% <strong>of</strong> total sum) and Quercoidites henrici (up<br />

to 6%), as well as Momipites punctatus, Castaneoideaepollis<br />

oviformis and C. pusillus (a few<br />

per cent). Pollen grains <strong>of</strong> palms (Arecipites<br />

pseudoconvexus) are encountered regularly,<br />

whereas Sapotaceae (Tetracolporopollenites<br />

andreanus and T. rotundus) and Meliaceae<br />

occur sporadically. In bottom samples <strong>of</strong> both<br />

pr<strong>of</strong>i les pollen grains <strong>of</strong> Reevesiapollis triangulus<br />

reach about 2%. Relatively high quantities<br />

<strong>of</strong> Tricolporopollenites exactus (up to<br />

20%), T. megaexactus (up to 1%), T. fallax (up<br />

to 12%), T. liblarensis (up to 3%), Araliaceaeoipollenites<br />

edmundi, A. euphorii (a few per<br />

cent), Ilexpollenites (I. iliacus, I. marcodurensis<br />

and I. propinquus – toge<strong>the</strong>r up to 10%), Myricipites<br />

+ Triatrio pollenites rurensis (up to 6%),<br />

Symplocoipollenites vestibulum and S. latiporis<br />

(toge<strong>the</strong>r up to 2%), as well as Corylopsis type<br />

(up to 1%) are also characteristic features <strong>of</strong><br />

this section. Moreover, pollen grains <strong>of</strong> Nelumbopollenites<br />

europaeus are sporadically found.<br />

Among <strong>the</strong> arctotertiary taxa <strong>the</strong> most<br />

numerous are Taxodiaceae/Cupressaceae (Inapertu<br />

ro pollenites – max. 50%), Nyssapollenites<br />

(up to 12%), Alnipollenites verus (a few per<br />

cent), Ericipites (up to 12%; including E. roboreus<br />

up to 2%), Sequoiapollenites (up to 35%),<br />

Pinuspollenites + Cathayapollis (up to 40%),<br />

and Abiespollenites (up to 12%). Pollen grains<br />

<strong>of</strong> Sciadopityspollenites are encountered regularly.<br />

In addition, in bottom samples Intratriporopollenites<br />

insculptus (present in only one<br />

sample) and Periporopollenites orien taliformis<br />

occur.<br />

Section L1 is characterized also by high frequency<br />

<strong>of</strong> Sphagnum spores (max. 35%), while<br />

Polypodiaceae s.l. and Osmunda, as well as<br />

spores <strong>of</strong> Lycopodiaceae and Toroisporis are<br />

found sporadically. In addition, <strong>the</strong> Microthyriaceae<br />

fungi, fragments <strong>of</strong> plant tissues as<br />

well as stomata resembling those <strong>of</strong> <strong>the</strong> Taxodiaceae<br />

family, and fresh-water phytoplankton<br />

(Ovoidites ligneolus, Tetraporina and Sigmopollis)<br />

were found. In bottom samples <strong>of</strong> both<br />

<strong>Legnica</strong> pr<strong>of</strong>i les Sigmopollis is very numerous,<br />

and accompanied by single din<strong>of</strong>l agellate<br />

cysts.<br />

In <strong>the</strong> upper part <strong>of</strong> this section <strong>of</strong> <strong>the</strong> pr<strong>of</strong>i le<br />

<strong>Legnica</strong> 33/56 frequency <strong>of</strong> Cyrillaceae/Clethraceae,<br />

Ericaceae, Ilex, Myrica, and Sequoia distinctly<br />

increases. Moreover, in <strong>the</strong>se samples<br />

<strong>the</strong> role <strong>of</strong> mesophilous taxa Tricolporopollenites<br />

pseudocingulum, Araliaceae, Fabaceae,

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