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Middle Miocene palynoflora of the Legnica lignite deposit complex ...

Middle Miocene palynoflora of the Legnica lignite deposit complex ...

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R e m a r k s. Pollen grains 45–55 × 40–45 μm<br />

in size, not numerous in <strong>the</strong> studied material.<br />

Pollen grains <strong>of</strong> fossil taxon Nyssapollenites<br />

kruschi (and its subspecies) occur in<br />

Europe in <strong>the</strong> Oligocene to Pliocene <strong>deposit</strong>s,<br />

and represent warm-temperate (A1) element<br />

(Ziembińska-Tworzydło 1996).<br />

(154) Nyssapollenites pseudocruciatus<br />

(Potonié 1931) Thiergart 1937<br />

Pl. 16, fi gs 7a, b, 8<br />

1931a Pollenites pseudocruciatus n. sp., Potonié,<br />

p. 328, pl. 1, fi g. 10.<br />

1937 Nyssapollenites pseudocruciatus (Potonié) n.<br />

comb., Thiergart, p. 322, pl. 25, fi gs 32–34,<br />

pl. 26, fi g. 1.<br />

Pollen grains 27–32 μm in size, resembling<br />

those <strong>of</strong> Nyssapollenites kruschi. Pores with<br />

very thick edges, surface granulate, somewhat<br />

more delicate than N. analepticus.<br />

R e m a r k s. This taxon represents warm-temperate<br />

(A1) element (Ziembińska-Tworzydło<br />

et al. 1994a). In <strong>the</strong> studied material <strong>the</strong>se<br />

pollen grains were encountered regularly, but<br />

in low quantities.<br />

Pollen grains <strong>of</strong> Nyssapollenites morphological<br />

genus are similar to pollen <strong>of</strong> Nyssa.<br />

They are frequent in <strong>the</strong> Neogene, though <strong>the</strong>y<br />

are reported also from <strong>the</strong> Palaeogene (Muller<br />

1981). In <strong>the</strong> younger Neogene larger forms<br />

prevail (Doktorowicz-Hrebnicka 1954).<br />

Presently <strong>the</strong> genus Nyssa contains 6 species<br />

<strong>of</strong> deciduous trees; four <strong>of</strong> <strong>the</strong>m occur in<br />

<strong>the</strong> Atlantic zone <strong>of</strong> North America, whereas<br />

<strong>the</strong> o<strong>the</strong>r two grow in south-eastern Asia<br />

(Krüssmann 1977). Nyssa aquatica L. and N.<br />

sylvatica Marsh are components <strong>of</strong> swamp forests<br />

(Kearney 1901, Kac 1975, Haynes 2000).<br />

Familia CORNACEAE<br />

Subfamilia CORNOIDEAE<br />

Cornaceaepollis Stuchlik in Ziembińska-<br />

Tworzydło et al. 1994<br />

(155) Cornaceaepollis major (Stuchlik<br />

1964) Stuchlik in Ziembińska-Tworzydło<br />

et al. 1994<br />

Pl. 16, fi g. 13a, b<br />

1964 Cornoidites major n. sp., Stuchlik, p. 62, pl. 19,<br />

fi gs 1–4.<br />

1994b Cornaceaepollis major (Stuchlik) Stuchlik comb.<br />

53<br />

nov.; Ziembińska-Tworzydło et al., p. 22, pl. 13,<br />

fi g. 11a–c.<br />

Pollen grains tricolporate, in equatorial<br />

view broadly oval in outline, 55–60 μm in<br />

size, in polar view triangularly oval in outline.<br />

Colpi ra<strong>the</strong>r long, bent in equatorial plain,<br />

nearly meeting each o<strong>the</strong>r at <strong>the</strong> poles. Pores<br />

in <strong>the</strong> middle <strong>of</strong> <strong>the</strong> colpi, set transversally to<br />

<strong>the</strong>m, up to 15 μm in diameter. Exine 4–5 μm<br />

thick. Ectexine thicker than endexine. Surface<br />

granulate.<br />

R e m a r k s. In <strong>the</strong> analysed material a few<br />

pollen grains <strong>of</strong> this taxon were found in <strong>the</strong><br />

Lusatian seam.<br />

(156) Cornaceaepollis minor (Stuchlik<br />

1964) Stuchlik in Ziembińska-Tworzydło<br />

et al. 1994<br />

Pl. 16, fi g. 11a, b<br />

1964 Cornoidites minor n. sp.; Stuchlik, p. 62, pl. 19,<br />

fi gs 5–7.<br />

1994b Cornaceaepollis minor (Stuchlik) Stuchlik comb.<br />

nov.; Ziembińska-Tworzydło et al., p. 22, pl. 13,<br />

fi gs 12a, b, 13.<br />

Pollen grains resembling in structure abovementioned<br />

ones, but 24–28 μm in size. Exine<br />

about 2 μm thick, surface fi nely granulate.<br />

R e m a r k s. Pollen grains <strong>of</strong> this taxon were<br />

sporadically encountered in <strong>the</strong> Lusatian seam<br />

<strong>of</strong> both <strong>Legnica</strong> pr<strong>of</strong>i les.<br />

Pollen grains <strong>of</strong> both above-mentioned taxa<br />

(Cornaceaepollis major and C. minor) are<br />

similar to pollen <strong>of</strong> <strong>the</strong> recent Cornus L. They<br />

occur in <strong>the</strong> Oligocene and <strong>Miocene</strong> <strong>deposit</strong>s<br />

<strong>of</strong> <strong>the</strong> Polish Lowland, and represent subtropical<br />

(P2) element (Ziembińska-Tworzydło et al.<br />

1994a, b).<br />

Nowadays <strong>the</strong> family Cornaceae contains 12<br />

genera extended in tropical to temperate zones<br />

<strong>of</strong> North America and in temperate Asia. The<br />

genus Cornus (about 40 species) is common<br />

throughout nor<strong>the</strong>rn temperate zone (Krüssmann<br />

1976).<br />

Subfamilia MASTIXIOIDEAE<br />

(157) Cornaceaepollis satzveyensis (Pfl ug<br />

in Thomson & Pfl ug 1953) Ziembińska-<br />

Tworzydło in Ziembińska-Tworzydło et al. 1994<br />

Pl. 16, fi g. 9a, b<br />

1953 Tricolporopollenites satzveyensis n. sp. (Pfl ug);<br />

Thomson & Pfl ug, p. 103, pl. 13, fi gs 10–13.

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