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Middle Miocene palynoflora of the Legnica lignite deposit complex ...

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cit.) confi rmed <strong>the</strong>ir botanical affi nity with<br />

<strong>the</strong> genus Reevesia. The morphological species<br />

Reevesiapollis triangulus represents subtropical<br />

(P2) element (Ziembińska-Tworzydło et al.<br />

1994a). It is characteristic for <strong>the</strong> <strong>Middle</strong> <strong>Miocene</strong><br />

to Upper Pliocene <strong>deposit</strong>s (Sadowska<br />

1973, Jahn et al. 1984). Tertiary localities <strong>of</strong><br />

Reevesia are known from sou<strong>the</strong>rn and central<br />

Europe, as well as from Asia (Petrov & Drazheva-Stamatova<br />

1972). In <strong>the</strong> studied material<br />

pollen grains <strong>of</strong> Reevesiapollis triangulus<br />

were encountered regularly, but in quantities<br />

not exceeding 1% (exceptionally reaching 2–3%<br />

in bottom samples <strong>of</strong> both <strong>Legnica</strong> pr<strong>of</strong>i les).<br />

Nowadays members <strong>of</strong> <strong>the</strong> genus Reevesia<br />

(15 species <strong>of</strong> trees and shrubs) are distributed<br />

in East Asia, from <strong>the</strong> Himalayas to Taiwan<br />

in evergreen broad-leaved forests and rain forests<br />

(Krüssmann 1978).<br />

Familiae STERCULIOIDEAE,<br />

RUTACEAE<br />

(123) Tricolporopollenites sp. 2<br />

Pl. 14, fi g. 12a, b.<br />

1993 Sterculiaceae–Rutaceae, Tricolporopollenites sp.<br />

12; Kohlman-Adamska, p. 149, pl. 25, fi g. 3a, b.<br />

Pollen grains tricolporate, in equatorial<br />

view circularly oval in outline, 22 × 18 μm in<br />

size. Colpi long, pores about 2 μm in diameter.<br />

Exine about 1 μm thick, surface reticulate<br />

with regular lumina (about 1 μm in diameter)<br />

and thin muri.<br />

R e m a r k s. A few pollen grains <strong>of</strong> this type <strong>of</strong><br />

structure were found in <strong>the</strong> analysed material.<br />

Ordo ROSALES<br />

Familia ULMACEAE<br />

Celtis L.<br />

Celtipollenites Nagy 1969 emend. Kohlman-<br />

Adamska & Ziembińska-Tworzydło<br />

in Stuchlik et al. 2009<br />

(124) Celtipollenites bobrowskae Kohlman-<br />

Adamska & Ziembińska-Tworzydło<br />

in Stuchlik et al. 2009<br />

Pl. 14, fi g. 18<br />

2009 Celtipollenites bobrowskae Kohlman-Adamska<br />

& Ziembińska-Tworzydło sp. nov.; Stuchlik et al.,<br />

p. 64, pl. 50, fi gs 10–15, pl. 51, fi gs 1–5.<br />

45<br />

R e m a r k s. These pollen grains are close to <strong>the</strong><br />

recent species Celtis sinensis Pers. Celtipollenites<br />

bobrowskae represents warm-temperate<br />

(A1) element, and is known from <strong>the</strong> <strong>Miocene</strong> –<br />

Pliocene <strong>deposit</strong>s (Stuchlik et al. 2009). These<br />

pollen grains were regularly encountered in<br />

all studied pr<strong>of</strong>i les in quantities not exceeding<br />

1–2%, only in <strong>the</strong> grey clay horizon reaching<br />

2–5%.<br />

(125) Celtipollenites komloensis Nagy 1969<br />

Pl. 14, fi g. 17<br />

1969 Celtipollenites komlóensis n. g. n. sp., Nagy,<br />

p. 456, pl. 43, fi gs 3, 7.<br />

1985 Celtipollenites komloënsis Nagy, p. 197, pl. 111,<br />

fi gs 17–20.<br />

2009 Celtipollenites komloensis Nagy; Stuchlik et al.,<br />

p. 65, pl. 50, fi gs 1–3, 5.<br />

R e m a r k s. These pollen grains are close to<br />

<strong>the</strong> recent species Celtis occidentalis L., and<br />

represent warm-temperate element (A1). They<br />

are reported from <strong>the</strong> <strong>Miocene</strong>-Pliocene <strong>deposit</strong>s.<br />

In Poland <strong>the</strong>y are rarely encountered in<br />

<strong>the</strong> <strong>Middle</strong> <strong>Miocene</strong> <strong>deposit</strong>s (Stuchlik et al.<br />

2009). Several pollen grains <strong>of</strong> this species<br />

were found in <strong>the</strong> studied material.<br />

The present-day genus Celtis contains about<br />

80 (100) species <strong>of</strong> trees, shrubs, rarely climbers,<br />

extended in temperate and tropical zones.<br />

Celtis sinensis Pers. is distributed in eastern<br />

China, Japan and Korea. Celtis occidentalis<br />

grows on humid soils in <strong>the</strong> Atlantic zone <strong>of</strong><br />

North America (Kearney 1901, Krüssmann<br />

1976, Kubitzki 1993).<br />

Ulmus L.<br />

Ulmipollenites Wolff 1934<br />

(126) Ulmipollenites maculosus Nagy 1969<br />

Pl. 14, fi g. 19<br />

1969 Ulmipollenites maculosus n. sp., Nagy, p. 223,<br />

pl. 52, fi gs 1, 2.<br />

R e m a r k s. These pollen grains represent<br />

cool-temperate element (A2), and occur in <strong>the</strong><br />

Oligocene to Pliocene <strong>deposit</strong>s (Ziembińska-<br />

Tworzydło 1996, Stuchlik et al. 2009). In <strong>the</strong><br />

studied material <strong>the</strong>se pollen grains occurred<br />

regularly in quantities 1–2%, in several<br />

samples <strong>of</strong> <strong>the</strong> grey clay horizon and Mużaków<br />

series reaching about 5%.

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