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Middle Miocene palynoflora of the Legnica lignite deposit complex ...

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44<br />

1953 Intratriporopollenites instructus (Potonié) n.<br />

comb., Thomson & Pfl ug, p. 89, pl. 10, fi gs<br />

10–23.<br />

R e m a r k s. According to many authors (Thomson<br />

& Pfl ug 1953, Thiele-Pfeiffer 1980, Planderová<br />

1990, Ziembińska-Tworzydło 1996) pollen<br />

grains <strong>of</strong> <strong>the</strong> morphological taxon Intratriporopollenites<br />

instructus resemble pollen <strong>of</strong> <strong>the</strong><br />

recent genus Tilia L. However, in <strong>the</strong> <strong>Miocene</strong><br />

<strong>deposit</strong>s <strong>of</strong> Wiesa (Germany) similar ones have<br />

been found toge<strong>the</strong>r with fl owers <strong>of</strong> <strong>the</strong> fossil<br />

species Burretia instructa (Potonié) Mai (Mai<br />

1961). According to this author <strong>the</strong> fossil fl owers<br />

resemble those <strong>of</strong> <strong>the</strong> recent genera Brownlowia<br />

Roxb. and Pentace Hassk. Pollen grains<br />

<strong>of</strong> Intratriporopollenites instructus occur in<br />

Europe in <strong>the</strong> Oligocene to Pliocene <strong>deposit</strong>s,<br />

with maximum in <strong>the</strong> <strong>Middle</strong> <strong>Miocene</strong>, and<br />

represent warm-temperate (A1) element (Mai<br />

1961, Ziembińska-Tworzydło 1996). Several<br />

pollen grains <strong>of</strong> this taxon were found in <strong>the</strong><br />

studied material.<br />

(120) Intratriporopollenites insculptus<br />

Mai 1961<br />

Pl. 14, fi gs 10, 11<br />

1961 Intratriporopollenites insculptus n. sp., Mai,<br />

p. 65, pl. 11, fi gs 10–27.<br />

R e m a r k s. This taxon is similar to pollen<br />

grains <strong>of</strong> <strong>the</strong> recent subfamily Brownlowioideae<br />

(Mai 1961). Brownlowia argentata is<br />

a component <strong>of</strong> mangrove formation on Borneo,<br />

where pollen grains <strong>of</strong> this type occur in <strong>deposit</strong>s<br />

since <strong>the</strong> Eocene (Muller 1964). In Poland<br />

Intra triporopollenites insculptus is known<br />

from <strong>the</strong> <strong>Miocene</strong> <strong>of</strong> central and south-western<br />

part <strong>of</strong> <strong>the</strong> Polish Lowland. They also occur in<br />

<strong>the</strong> <strong>Miocene</strong> <strong>of</strong> Hungary, Oligocene – <strong>Miocene</strong><br />

<strong>of</strong> Germany and former Soviet Union, as well<br />

Pliocene <strong>of</strong> Slovakia (Ziembińska-Tworzydło<br />

1996). They represent warm-temperate (A1)<br />

element (Ziembińska-Tworzydło 1996). Only 3<br />

pollen grains <strong>of</strong> this taxon were found in bottom<br />

sample <strong>of</strong> <strong>Legnica</strong> 33/56 pr<strong>of</strong>i le.<br />

Pollen grains <strong>of</strong> <strong>the</strong> two above-mentioned<br />

morphological species (Intratriporopollenites<br />

insculptus and I. instructus) were isolated<br />

from fl ower buds <strong>of</strong> one fossil species Craigia<br />

bronnii (Unger) Z. Kvaček, Bůžek et Manchester,<br />

from <strong>the</strong> Late <strong>Miocene</strong> fl ora <strong>of</strong> Hambach,<br />

Germany (Kvaček et al. 2002).<br />

Subfamilia TILIOIDEAE<br />

(121) Intratriporopollenites<br />

cordataeformis (Wolff 1934) Mai 1961<br />

Pl. 14, fi g. 8a, b<br />

1934 Tiliae-pollenites instructus cordataeformis n. f.,<br />

Wolff, p. 73, pl. 5, fi g. 22.<br />

1961 Intratriporopollenites cordataeformis (Wolff) n.<br />

comb., Mai, p. 67, pl. 13, fi gs 4–7.<br />

R e m a r k s. These pollen grains are <strong>the</strong> nearest<br />

pollen <strong>of</strong> recent Tilia cordata Mill., and represent<br />

cool-temperate (A2) element (Ziembińska-<br />

Tworzydło et al. 1994a). According to Mai<br />

(1961) Intratriporopollenites cordataeformis is<br />

a Pliocene taxon. Similar pollen grains have<br />

been found in <strong>the</strong> Upper <strong>Miocene</strong> and Pliocene<br />

<strong>of</strong> Paratethys (Planderová 1990), and <strong>Middle</strong><br />

<strong>Miocene</strong> <strong>of</strong> Sudetic Foredeep (Ziembińska-<br />

Tworzydło et al. 1994a). Only 3 pollen grains <strong>of</strong><br />

this type were found in <strong>the</strong> analysed material<br />

in one sample from <strong>the</strong> Mużaków series.<br />

Nowadays <strong>the</strong> former family Tiliaceae contains<br />

about 41 genera and 400 species <strong>of</strong> trees<br />

and shrubs, rarely herbs, extended mainly in<br />

temperate zone. The genus Tilia (about 30<br />

species <strong>of</strong> deciduous trees) occurs in nor<strong>the</strong>rn<br />

temperate zone (Heywood 1978, Bugała 1991).<br />

According to Mai (1961) <strong>the</strong> Tilia genus can<br />

be demonstrated with certainty by pollen only<br />

since <strong>the</strong> Pliocene.<br />

Subfamilia STERCULIOIDEAE<br />

Reevesia Lindl.<br />

Reevesiapollis Krutzsch 1970<br />

(122) Reevesiapollis triangulus<br />

(Mamczar 1960) Krutzsch 1970<br />

Pl. 14, fi g. 13<br />

1960 Pollenites triangulus n. spm., Mamczar, p. 220,<br />

pl. 14, fi g. 202.<br />

1970c Reevesiapollis triangulus (Mamczar) comb.<br />

nov., Krutzsch, p. 374, pl. 5, fi gs 19–35, pl. 6,<br />

fi gs 1–11, pl. 7, fi gs 1–44, pl. 8, fi gs 1–21.<br />

R e m a r k s. Pollen grains <strong>of</strong> similar structure<br />

have been previously described under<br />

various names (see Sadowska 1973), and connected<br />

with <strong>the</strong> recent families Salicaceae,<br />

Oleaceae, Caprifoliaceae, Hamamelidaceae,<br />

Tiliaceae, and Balsaminaceae. Sadowska (op.

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