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Middle Miocene palynoflora of the Legnica lignite deposit complex ...

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38<br />

1994b Corylopollis coryloides (Pfl ug) Ziembińska-<br />

Tworzydło comb. nov.; Ziembińska-Tworzydło<br />

et al., p. 16, pl. 8, fi gs 18, 19.<br />

R e m a r k s. These pollen grains represent<br />

arctotertiary element (A). They occur in fossil<br />

palyn<strong>of</strong>l oras since <strong>the</strong> Oligocene, more frequently<br />

in <strong>the</strong> Neogene, and commonly in <strong>the</strong><br />

Pliocene (Stuchlik et al. 2009). Pollen grains<br />

<strong>of</strong> this taxon were encountered in all studied<br />

pr<strong>of</strong>i les, mainly in <strong>the</strong> grey clay horizon and<br />

Henryk seam, in quantities not exceeding 1%.<br />

Today Corylus genus (about 15–19 species<br />

<strong>of</strong> tall shrubs and trees) occurs in nor<strong>the</strong>rn<br />

temperate zone <strong>of</strong> Europe, America and Asia<br />

(Krüssmann 1976, Kubitzki 1993).<br />

Betula L.<br />

Trivestibulopollenites Pfl ug in Thomson<br />

& Pfl ug 1953<br />

(96) Trivestibulopollenites betuloides<br />

Pfl ug in Thomson & Pfl ug 1953<br />

Pl. 12, fi g. 8<br />

1953 Trivestibulopollenites betuloides n. sp. (Pfl ug);<br />

Thomson & Pfl ug, p. 85, pl. 9, fi gs 25–34.<br />

1969 Betulaepollenites betuloides (Pfl ug) n. comb.,<br />

Nagy, p. 228, pl. 3, fi g. 12.<br />

R e m a r k s. Pollen grains <strong>of</strong> Betula type occur<br />

in Europe since <strong>the</strong> Palaeocene ( Krutzsch<br />

1970d, Muller 1981). Trivestibulopollenites<br />

betuloides occurs in <strong>the</strong> Eocene to Pliocene<br />

<strong>deposit</strong>s, and represents arctotertiary (A) element<br />

(Ziembińska-Tworzydło et al. 1994a,<br />

Stuchlik et al. 2009). Pollen grains <strong>of</strong> T. betuloides<br />

were regularly noted in all analysed<br />

pr<strong>of</strong>i les in quantities <strong>of</strong> a few per cent (max.<br />

10–12% in <strong>the</strong> Mużaków series and Lusatian<br />

seam).<br />

The recent species <strong>of</strong> Betula genus (about<br />

60) are distributed in temperate and boreal<br />

zones <strong>of</strong> <strong>the</strong> nor<strong>the</strong>rn hemisphere (Krüssmann<br />

1976). They are photophilous plants, but <strong>the</strong>y<br />

have little requirements about soil (Bugała<br />

1991).<br />

Familia MYRICACEAE<br />

Myrica L.<br />

Myricipites Wodehouse 1933 emend.<br />

Grabowska & Ważyńska<br />

in Stuchlik et al. 2009<br />

(97) Myricipites pseudorurensis (Pfl ug<br />

in Thomson & Pfl ug 1953) Grabowska<br />

& Ważyńska in Stuchlik et al. 2009<br />

Pl. 12, fi g. 17<br />

1953 Triatriopollenites pseudorurensis n. sp. (Pfl ug);<br />

Thomson & Pfl ug, p. 79, pl. 7, fi gs 76–80.<br />

2009 Myricipites pseudorurensis (Pfl ug in Thomson<br />

& Pfl ug) Grabowska & Ważyńska comb. nov.;<br />

Stuchlik et al., p. 27, pl. 16, fi gs 1–7.<br />

R e m a r k s. These pollen grains resemble<br />

recent pollen <strong>of</strong> Myrica gale L. They occur in<br />

<strong>the</strong> Eocene to Pliocene, and represent arctotertiary<br />

(A) element. In Poland <strong>the</strong>y are reported<br />

from <strong>the</strong> Upper Oligocene to <strong>Miocene</strong> <strong>deposit</strong>s<br />

(Stuchlik et al. 2009). They were regularly<br />

encountered in <strong>the</strong> studied material.<br />

Triatriopollenites Pfl ug & Thomson<br />

in Thomson & Pfl ug 1953<br />

(98) Triatriopollenites rurensis Pfl ug<br />

& Thomson in Thomson & Pfl ug 1953<br />

Pl. 15, fi gs 18, 19<br />

1953 Triatriopollenites rurensis n. sp. (Pfl ug & Thomson);<br />

Thomson & Pfl ug, p. 79, pl. 7, fi gs 81–109.<br />

1969 Myricipites rurensis (Pfl ug & Thomson) n. comb.,<br />

Nagy, p. 245, pl. 53, fi g. 27.<br />

R e m a r k s. These pollen grains represent subtropical/arctotertiary<br />

(P2/A) element, and occur<br />

in Poland in <strong>the</strong> Eocene to <strong>Miocene</strong> <strong>deposit</strong>s<br />

(Stuchlik et al. 2009). Kohlman-Adamska<br />

(1993) compares this species with Myrica nagy<br />

Thunb. and M. salicifolia Hochst. ex Rich. Pollen<br />

grains <strong>of</strong> Triatriopollenites rurensis were<br />

regularly encountered in <strong>the</strong> studied material.<br />

They were more frequent in <strong>the</strong> Lusatian seam.<br />

The oldest pollen grains <strong>of</strong> Myrica type<br />

are known from <strong>the</strong> Cretaceous (Santonian)<br />

<strong>of</strong> USA. In Europe <strong>the</strong>y are more frequent<br />

since <strong>the</strong> Eocene and Oligocene (Muller 1981).<br />

Nowadays this genus (with about 35 species<br />

<strong>of</strong> evergreen or deciduous trees and shrubs) is<br />

distributed in temperate and subtropical zones<br />

<strong>of</strong> both hemispheres. Myrica adenophora grows<br />

in China, M. carolinensis in North America,<br />

M. nagy in north-western Himalayas, M. rubra<br />

in Asia, M. salicifolia in Ethiopia (Krüssmann<br />

1977). M. gale occurs on sea-shores <strong>of</strong> nor<strong>the</strong>rn<br />

and western Europe in maritime climate zone<br />

and in nor<strong>the</strong>rn part <strong>of</strong> North America. Morella<br />

cerifera (L.) Small. (=Myrica cerifera L.)

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