Middle Miocene palynoflora of the Legnica lignite deposit complex ...

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recent Quercus robur L. from Europe, North
America and Asia Minor. Quercoidites asper
represents warm-temperate element (A1),
in Europe it occurs in the Oligocene to Pliocene
deposits (Ziembińska-Tworzydło et al.
1994a, b, Ziembińska-Tworzydło 1996). These
pollen grains were regularly found in all studied
profi les.
(89) Quercoidites granulatus (Nagy 1969)
Słodkowska in Ziembińska-Tworzydło
et al. 1994
Pl. 12, fi g. 5
1969 Quercopollenites granulatus n. sp., Nagy, p. 223,
pl. 52, fi g. 21.
1994b Quercoidites granulatus (Nagy) Słodkowska
comb. nov.; Ziembińska-Tworzydło et al., p. 26,
pl. 15, fi gs 10, 11.
R e m a r k s. Pollen grains with coarser granulate
sculpture. The taxon Q. granulatus
represents warm-temperate element (A1), and
occurs in the Miocene deposits (Ziembińska-
Tworzydło et al 1994a, b). In the studied material
pollen grains of this taxon were found sporadically.
(90) Quercoidites henrici (Potonié 1931)
Potonié, Thomson & Thiergart 1950
Pl. 12, fi g. 6
1931a Pollenites henrici n. sp., Potonié, p. 332, pl. 2,
fi g. 19.
1950 Quercoidites henrici (Potonié); Potonié, Thomson
& Thiergart, p. 54, pl. B, fi gs 22, 23.
1953 Tricolpopollenites henrici (Potonié) n. comb.,
Thomson & Pfl ug, p. 95, p. 11, fi gs 39, 40.
1961 Tricolporopollenites henrici (Potonié) n. comb.,
Krutzsch, p. 322.
R e m a r k s. Pollen grains of this taxon were
often found in the Lusatian seam (up to 6%)
and Henryk seam (a few percent), in other
horizons they were encountered sporadically.
(91) Quercoidites microhenrici (Potonié
1931) Potonié, Thomson & Thiergart 1950
Pl. 12, fi g. 7
1931b Pollenites microhenrici n. sp., Potonié, p. 26.
1950 Quercoidites microhenrici (Potonié) n. comb.,
Potonié, Thomson & Thiergart, p. 55, pl. B, fi gs
24, 25, pl. C, fi g. 22.
1953 Tricolpopollenites microhenrici (Potonié) n.
comb., Thomson & Pfl ug, p. 95, p. 11, fi gs
62–110.
1961 Tricolporopollenites microhenrici (Potonié) n.
comb., Krutzsch, p. 322.
1994a Quercoidites microhenrici (Potonié) Potonié,
Thomson & Thiergart; Ziembińska-Tworzydło
et al., pl. 15, fi gs 5, 6.
R e m a r k s. Several pollen grains of this taxon
were found in the Lusatian seam.
The taxa Quercoidites henrici and Q. microhenrici
represent subtropical element (P2) and
occur in the Polish Lowland in the Upper Palaeocene
to Middle Miocene deposits, whereas in
Slovakia, Turkey and former Soviet Union to
the Pliocene (Grabowska 1996a).
The recent members of genus Quercus (about
350–450 species of deciduous and evergreen
trees and shrubs) are distributed in temperate
and subtropical zones of the northern hemisphere,
as well as in the mountains of tropical
Malaysia and north-western part of North
America (Krüssmann 1977, Kubitzki 1993).
Familia BETULACEAE
Alnus Mill.
Alnipollenites Potonié 1934
(92) Alnipollenites verus (Potonié 1931)
Potonié 1934
Pl. 12, fi gs 9, 10
1931a Pollenites verus n. sp., Potonié, p. 320, pl. 2,
fi g. 40.
1934 Alnipollenites verus (Potonié); Potonié, p. 58,
pl. 2, fi gs 13, 17, 18, 25, 26.
1953 Polyvestibulopollenites (Alnipollenites) verus
(Potonié) n. comb., Thomson & Pfl ug, p. 90,
pl. 10, fi gs 62–76.
R e m a r k s. These pollen grains correspond
to the recent pollen of Alnus Mill., especially
A. glutinosa (L.) Gaertn., A. hirsuta Turcz.,
A. incana (L.) Moench., A. rugosa (Du Roi)
Spreng., and A. tenuifolia Nutt. In previous
elaborations tetraporate grains 15–25 μm
in size were described as Alnus kefersteinii
Goepp. type, whereas bigger forms (above 30
μm in diameter), mainly 5- 6-porate as Alnus
glutinosa (L.) Gaertn. type (Doktorowicz-
Hrebnicka 1956a, Oszast 1960, 1973, Stuchlik
1964). Recently pollen grains of Alnipollenites
verus type were isolated from male catkins of
Alnus kefersteinii (Goeppert) Unger, found in
the Oligocene deposits at Bechlejovice, Czech
Republic (Dašková 2008). Alnipollenites verus