Middle Miocene palynoflora of the Legnica lignite deposit complex ...

1960 Rhoipites (Pollenites) pseudocingulum (Potonié)
n. comb., Potonié, p. 101, pl. 6, fi g. 114.
2000 Tricolporopollenites pseudocingulum (Potonié)
Thomson & Pfl ug (s.s.); Kohlman-Adamska
& Ziembińska-Tworzydło, p. 51, pl. 2, fi gs 2–4,
pl. 3, fi gs 1–5.
R e m a r k s. This taxon used to be connected
with the recent family Anacardiaceae and the
genera: Rhus (Tourn.) L., Mangifera L., and
Spondias L. (Mamczar 1962). However, detailed
studies of pollen grains under SEM revealed
their similarity to recent pollen of the family
Fagaceae, subfamily Fagoidae, and within
the collective taxon Tricolporopollenites pseudocingulum
(Potonié) Thomson & Pfl ug (s.l.)
a few species varying in morphology of tectum
sculpture have been distinguished (Kohlman-
Adamska & Ziembińska-Tworzydło 1999, 2000).
This taxon represents subtropical element (P2),
in Europe it occurs in the Palaeo gene to Upper
Miocene deposits (Ziembińska-Tworzydło 1996).
Pollen grains of T. pseudocingulum were very
often found in the studied material, in quantities
of a few per cent in the Henryk seam and
max. 42% in the Lusatian seam.
(86) Tricolporopollenites theacoides
(Roche & Schuler 1976) Kohlman-Adamska
& Ziembińska-Tworzydło 2000
Pl. 12, fi g. 3
1974 Rhoipites pseudocingulum (Potonié) Potonié;
Ziembińska-Tworzydło, pl. 23, fi g. 5.
1976 Verrutricolporites theacoides n. f sp., Roche
& Schuler, p. 27, pl. 10, fi gs 31, 32.
2000 Tricolporopollenites theacoides (Roche & Schuler)
comb. nov., Kohlman-Adamska & Ziembińska-
Tworzydło, p. 54, pl. 5, fi gs 3–5.
Pollen grains with coarsely verrucate tectum.
Verrucae 1.5–2.0 μm in diameter and 3–5
μm high.
R e m a r k s. Pollen grains of Tricolporopollenites
theacoides were sporadically encountered
in the studied material.
Fagus L.
Faguspollenites Raatz 1937 ex Potonié 1960
(87) Faguspollenites verus Raatz 1937
Pl. 11, fi g. 15
1937 Fagus-pollenites verus n. sp., Raatz, p. 23, pl. 1,
fi g. 17.
35
R e m a r k s. This fossil species represents
cool-temperate (A2) element (Ziembińska-
Tworzydło et al. 1994a). Within fossil pollen
of Fagus some authors distinguish two morphological
types: grains 30–35 μm in size with
coarser sculpture are reported as Fagus ferruginea
type, whereas bigger ones with delicate
sculpture – as F. orientalis type (Oszast
1960, Stuchlik 1964, Nagy 1969). Kohlman-
Adamska (1993) has also distinguished two
morphological types differing in sculpture and
thickness of exine: type 1 (F. sylvatica and
F. longipetiolata) and type 2 (F. orientalis and
F. ferruginea). Konzalová (1976) has recorded
Faguspollenites verus from the Lower Miocene
of Bohemia, but in Europe this taxon is more
frequent since the Middle Miocene, becoming
numerous in the Pliocene. In the studied material
pollen grains of F. verus were encountered
regularly, in both morphological types, in
quantities 2–4%.
The present-day genus Fagus, with about
10 species of trees, occurs in temperate zone of
the northern hemisphere. Fagus ferruginea Ait
occurs in North America, F. sylvatica L. in central
Europe to the Caucasus, F. orientalis Lipsky
grows on lower altitudes of the Caucasus,
F. longiopetiolata Seemen. – in mixed mesophytic
forests and evergreen broad-leaved forests
in central and south-eastern China (Wang
1961, Krüssmann 1977, Kubitzki 1993).
Subfamilia QUERCOIDEAE
Quercus L.
Quercoidites (Potonié, Thomson & Thiergart
1950 ex Potonié 1960) Słodkowska in
Ziembińska-Tworzydło et al. 1994
(88) Quercoidites asper (Pfl ug & Thomson
1953) Słodkowska in Ziembińska-Tworzydło
et al. 1994
Pl. 12, fi g. 4a, b
1953 Tricolpopollenites asper n. sp. (Pfl ug & Thomson);
Thomson & Pfl ug, p. 96, pl. 11, fi gs
43–47.
1961 Tricolporopollenites asper (Potonié) n. comb.,
Krutzsch, p. 322.
1994b Quercoidites asper (Pfl ug & Thomson) Słodkowska
comb. nov.; Ziembińska-Tworzydło et al.,
p. 26, pl. 15, fi g. 1.
R e m a r k s. This species approaches the