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Middle Miocene palynoflora of the Legnica lignite deposit complex ...

Middle Miocene palynoflora of the Legnica lignite deposit complex ...

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Familia ALTINGIACEAE<br />

Liquidambar L.<br />

Periporopollenites Pfl ug & Thomson<br />

in Thomson & Pfl ug 1953<br />

(78) Periporopollenites orientaliformis<br />

(Nagy 1969) Kohlman-Adamska<br />

& Ziembińska-Tworzydło<br />

in Stuchlik et al. 2009<br />

Pl. 11, fi gs 7a–c, 9 a, b<br />

1969 Liquidambarpollenites orientaliformis n. sp.,<br />

Nagy, p. 171, pl. 42, fi gs 1, 2.<br />

2009 Periporopollenites orientaliformis (Nagy) Kohlman-Adamska<br />

& Ziembińska-Tworzydło comb.<br />

nov.; Stuchlik et al., p. 71, pl. 58, fi gs 1–3, 6–11,<br />

pl. 59, fi gs 1–3.<br />

R e m a r k s. These pollen grains are similar<br />

to pollen <strong>of</strong> <strong>the</strong> recent Liquidambar orientalis<br />

L. from Asia Minor (Nagy 1969). They differ<br />

from Periporopollenites stigmosus in ellipsoidal<br />

pores. Periporopollenites orientaliformis represents<br />

warm-temperate (A1) element, and occurs<br />

in <strong>the</strong> <strong>Miocene</strong> – Pliocene <strong>deposit</strong>s. In Poland,<br />

pollen grains <strong>of</strong> this species are scattered in<br />

<strong>the</strong> <strong>Middle</strong> and Upper <strong>Miocene</strong> (Stuchlik et al.<br />

2009). They were rarely found in <strong>the</strong> studied<br />

material, mainly in <strong>the</strong> Lusatian seam.<br />

(79) Periporopollenites stigmosus (Potonié<br />

1931) Thomson & Pfl ug 1953<br />

Pl. 11, fi gs 6a, b, 8<br />

1931a Pollenites stigmosus n. sp., Potonié, p. 332, pl. 2,<br />

fi g. 1.<br />

1937 Liquidambarpollenites stigmosus (Potonié) n.<br />

comb., Raatz, p. 17, fi g. 26.<br />

1953 Periporopollenites stigmosus (Potonié) n. comb.,<br />

Thomson & Pfl ug, p. 111, pl. 15, fi g. 58.<br />

R e m a r k s. This species is morphologically<br />

identical with pollen <strong>of</strong> <strong>the</strong> recent Liquidambar.<br />

This is confi rmed by fi nding this type <strong>of</strong><br />

pollen in situ in fossil infl orescence <strong>of</strong> Liquidambar<br />

europea A. Braun (Kohlman-Adamska<br />

et al. 2004, Stuchlik et al. 2007). Periporopollenites<br />

stigmosus is most similar to pollen <strong>of</strong> <strong>the</strong><br />

recent Liquidambar styracifl ua L. from North<br />

America (Nagy 1969), which grows in mixed<br />

forests with Liriodendron tulipifera, Tsuga<br />

canadensis, and T. caroliniana, on humid<br />

soils. Periporopollenites stigmosus represents<br />

warm-temperate (A1) element, and occurs in<br />

<strong>the</strong> <strong>Miocene</strong> to Pliocene <strong>deposit</strong>s. In Poland it<br />

33<br />

is encountered in <strong>the</strong> <strong>Middle</strong> and Upper <strong>Miocene</strong><br />

(Stuchlik et al. 2009). Pollen grains <strong>of</strong><br />

this species were regularly encountered in <strong>the</strong><br />

studied pr<strong>of</strong>i les, particularly in <strong>the</strong> Lusatian<br />

seam, but in quantities not exceeding 2%.<br />

Pollen grains <strong>of</strong> Liquidambar type occur<br />

in fossil palyn<strong>of</strong>l oras since <strong>the</strong> Palaeocene,<br />

but <strong>the</strong>y are more frequent since <strong>the</strong> Eocene<br />

(Kuprianova 1960, Muller 1981).<br />

Today <strong>the</strong> genus Liquidambar (about 6<br />

species <strong>of</strong> trees) grows in East Asia (sou<strong>the</strong>rn<br />

China, Taiwan, Vietnam), eastern part <strong>of</strong><br />

<strong>the</strong> Mediterranean Region (Turkey, Rhodes,<br />

Cyprus), south-eastern part <strong>of</strong> North America,<br />

and Mexico and Nicaragua in central America<br />

(Krüssmann 1977, Kubitzki 1993).<br />

Familia HAMAMELIDACEAE<br />

Tricolporopollenites Pfl ug & Thomson<br />

in Thomson & Pfl ug 1953<br />

Corylopsis Sieb. & Zucc.<br />

(80) Tricolporopollenites sp. 1 –<br />

Corylopsis type sensu Oszast 1960<br />

Pl. 11, fi g. 12a, b<br />

1960 Corylopsis sp.; Oszast, p. 24, pl. 8, fi gs 6, 8–10.<br />

R e m a r k s. Pollen grains <strong>of</strong> this type occur in<br />

<strong>the</strong> Neogene sediments and are usually named<br />

Corylopsis (Oszast 1960, 1967, 1973, Stuchlik<br />

1964, Oszast & Stuchlik 1977, Skawińska<br />

1989). They were sporadically found in <strong>the</strong> studied<br />

material.<br />

The present-day genus Corylopsis includes<br />

about 20 (7–22) species <strong>of</strong> deciduous shrubs<br />

and trees, which grow from Assam to Japan,<br />

with <strong>the</strong> greatest variability in China (Krüssmann<br />

1976, Kubitzki 1993).<br />

Parrotia C.A. Mey., Distylium<br />

Sieb. & Zucc.<br />

(81) Tricolporopollenites indeterminatus<br />

(Romanowicz 1961)<br />

Ziembińska-Tworzydło 1974<br />

Pl. 11, fi g. 13<br />

1961 Pollenites indeterminatus n. sp., Romanowicz,<br />

p. 355, pl. 21, fi g. 275.<br />

1974 Tricolporopollenites indeterminatus (Romanowicz)<br />

n. comb., Ziembińska-Tworzydło, p. 397,<br />

pl. 24, fi g. 9a, b.<br />

R e m a r k s. Pollen grains <strong>of</strong> this species

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