Middle Miocene palynoflora of the Legnica lignite deposit complex ...

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Keteleeria Carr.
Keteleeriapollenites Nagy 1969
(52) Keteleeriapollenites dubius
(Khlonova 1960) Słodkowska
in Ziembińska-Tworzydło et al. 1994
Pl. 6, fi g. 3
1960 Keteleeria dubia n. sp., Khlonova, p. 59, pl. 9,
fi g. 5.
1971 Abiespollenites dubius (Khlonova) n. comb.,
Krutzsch, p. 98, pl. 21.
1994b Keteleeriapollenites dubius (Khlonova)
Słodkowska comb. nov.; Ziembińska-Tworzydło
et al., p. 14, pl. 7, fi g. 4.
R e m a r k s. Pollen grains distinctly differing
from those of Abies in the lack of crest
on their corpus; connected with the genus
Keteleeria, and warm-temperate (A1) element
(Ziembińska-Tworzydło et al. 1994a,
b). Macro-remains and fossil pollen grains
of Keteleeria are known from almost whole
Europe, from the Upper Oligocene to Pliocene
localities (Khlonova 1960, Krutzsch 1971,
Ziembińska-Tworzydło 1974, Nagy 1985,
Kohlman-Adamska 1993, Uzunova & Ivanov
1996). In Russia they are reported from the
Upper Cretaceous to Pliocene, whereas in
Poland from the Lower Miocene to Plio cene
deposits (Ziembińska-Tworzydło 1996, Stuchlik
et al. 2002). Uzunova and Ivanov (1996)
consider Keteleeria to be a component of the
Tertiary mesophytic forests in the mountains
and, according to these authors, migrations of
these trees refl ect palaeoclimate fl uctuations.
In the studied material pollen grains of Keteleeriapollenites
dubius were found sporadically.
Presently the genus Keteleeria (4–8 species
of trees) is distributed in China, Taiwan,
Vietnam and Laos (Krüssmann 1972, Stuchlik
et al. 2002).
Larix Mill.
Laricispollenites Nagy 1985
(53) Laricispollenites sp.
Pl. 6, fi g. 4
Pollen grains inaperturate, circular in outline,
70–80 μm in diameter. Exine about 1 μm
thick, surface psilate.
R e m a r k s. Similar forms were described as
Larix sp., Phohosphaera pseudotsugoides, Pseudotsuga
sp., Laricoidites magnus, cf. Laevigatosporites
sp., Inaperturopollenites magnus, and
others (Krutzsch 1971, Ziembińska-Tworzydło
1974). Pollen grains of Laricispollenites genus
occur in the Miocene deposits, and represent
arctotertiary (A) element (Stuchlik et al.
2002). Only two pollen grains of Laricispollenites
sp. were found in the Mużaków series
and Lusatian seam.
Today the genus Larix (about 10 species)
is distributed in cooler areas of the northern
hemisphere. The genus Pseudotsuga contains
about 20 species of evergreen trees extended
in East Asia and North America (Krüssmann
1972).
Picea A. Dietr.
Piceapollis Krutzsch 1971
(54) Piceapollis planoides Krutzsch 1971
ex Hochuli 1978
Pl. 6, fi g. 5
1971 Piceapollis planoides n. sp. (=?Picea planoides n.
sp.), Krutzsch, p. 110, pl. 25, fi gs 1–4.
1978 Piceapollis planoides Krutzsch; Hochuli, p. 67,
pl. 8, fi g. 2.
1985 Piceapollenites planoides (Krutzsch) n. comb.,
Nagy, p. 138, pl. 69, fi g. 1.
R e m a r k s. Pollen grains of this taxon resemble
pollen of the recent Picea. They occur in
the Lower Miocene to Lower Pliocene deposits,
and represent warm-temperate (A1) element
(Stuchlik et al. 2002). Several pollen grains
of this taxon were found in the studied material.
(55) Piceapollis sacculiferoides Krutzsch
1971 ex Hochuli 1978
Pl. 7, fi g. 1
1971 Piceapollis sacculiferoides n. sp. (=?Picea sacculiferoides
n. sp.), Krutzsch, p. 108, pl. 24, fi gs
1–8.
1978 Piceapollis sacculiferoides Krutzsch; Hochuli,
p. 67, pl. 7, fi g. 10.
1985 Piceapollenites sacculiferoides (Krutzsch)
n. comb., Nagy, p.138, pl. 69, fi gs 2–5.
R e m a r k s. Pollen grains of Picea type, having
no living relative species equivalent. They
occur in the Miocene and Pliocene deposits
(Krutzsch 1971, Nagy 1985), and represent