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Middle Miocene palynoflora of the Legnica lignite deposit complex ...

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24<br />

1974 Abiespollenites latisaccatus (Trevisan) Krutzsch;<br />

Ziembińska-Tworzydło, p. 348, p. 10, fi g. 1.<br />

R e m a r k s. Pollen grains similar to pollen <strong>of</strong><br />

Abies fi rma Sieb. & Zucc., A. veitchii Lindl.,<br />

A. nordmanniana (Stev.) Spach, as well Pinus<br />

taeda L. They represent arctotertiary (A) element,<br />

and occur in fossil micr<strong>of</strong>l oras since<br />

<strong>the</strong> Eocene/Oligocene boundary, mainly in<br />

cooler climatic phases, being more frequent<br />

since <strong>the</strong> Pliocene. They are also known from<br />

<strong>the</strong> Polish <strong>Middle</strong> <strong>Miocene</strong> to Pliocene localities<br />

(Ziembińska-Tworzydło 1974, Kohlman-<br />

Adamska 1993, Ziembińska-Tworzydło et al.<br />

1994a, Stuchlik et al. 2002). In <strong>the</strong> analysed<br />

material <strong>the</strong>y were encountered regularly, in<br />

quantities <strong>of</strong> a few per cent.<br />

(45) Abiespollenites maximus Krutzsch<br />

1971 ex Nagy 1985<br />

Pl. 4, fi g. 11<br />

1971 Abiespollenites maximus n. sp., Krutzsch p. 92,<br />

pl. 18.<br />

1985 Abiespollenites maximus Krutzsch; Nagy, p. 139,<br />

pl. 72, fi g. 1.<br />

R e m a r k s. According to Krutzsch (1971)<br />

large dimensions <strong>of</strong> <strong>the</strong> pollen grains (150–170<br />

μm) suggest <strong>the</strong>ir affi nity with recent genus<br />

Keteleeria. However, <strong>the</strong> presence <strong>of</strong> a thick<br />

crest on <strong>the</strong>ir corpus supports <strong>the</strong>ir affi nity<br />

to Abies. Pollen grains <strong>of</strong> this fossil taxon are<br />

known from <strong>the</strong> Lower <strong>Miocene</strong> to Pliocene. In<br />

Poland <strong>the</strong>y occur in Lower and <strong>Middle</strong> <strong>Miocene</strong><br />

palyn<strong>of</strong>l oras. This taxon is included into<br />

arctoter tiary (A) element (Stuchlik et al. 2002).<br />

Several pollen grains <strong>of</strong> Abiespollenites maximus<br />

were found in <strong>the</strong> analysed material.<br />

Nowadays <strong>the</strong> genus Abies (about 50 species)<br />

is common in <strong>the</strong> nor<strong>the</strong>rn hemisphere,<br />

mostly in East Asia (China and Japan) and<br />

western North America. These trees grow<br />

mainly in <strong>the</strong> mountains, <strong>of</strong>ten on high altitudes;<br />

several lowland species have extensive<br />

ranges, but most <strong>of</strong> <strong>the</strong>m occur locally. Their<br />

climatic and soil requirements depend on <strong>the</strong><br />

species (Bugała 1991). Abies alba is distributed<br />

in <strong>the</strong> mountains <strong>of</strong> sou<strong>the</strong>rn, western and<br />

central Europe, Caucasus, Ukraine, and Belarus.<br />

Abies fi rma and A. veitchii occur in Japan,<br />

A. nordmanniana grows in western Caucasus<br />

and nor<strong>the</strong>rn Turkey (Stuchlik et al. 2002).<br />

Cathaya Chun & Kuang<br />

Cathayapollis Ziembińska-Tworzydło<br />

in Stuchlik et al. 2002<br />

Sivak (1976) among fossil pollen grains <strong>of</strong><br />

Cathaya type from France distinguished 12<br />

species differing in shape, and grouped <strong>the</strong>m<br />

into six morphological types <strong>of</strong> <strong>the</strong> sacci attachment<br />

as well as surface <strong>of</strong> corpus and sacci (see<br />

Stuchlik et al. 2002).<br />

(46) Cathayapollis erdtmanii (Sivak 1976)<br />

Ziembińska-Tworzydło in Stuchlik et al. 2002<br />

Pl. 5, fi g. 1a, b<br />

1949 Pinus haploxylon – Form Rudolph; Kremp, p. 60,<br />

pl. 4, fi g. 24.<br />

1976 Cathaya erdtmanii sp. nov., Sivak, p. 264, pl. 5,<br />

fi gs 1, 5, pl. 6. fi gs 1–7.<br />

2002 Cathayapollis erdtmanii (Sivak) Ziembińska-<br />

Tworzydło comb. nov.; Stuchlik et al., p. 15,<br />

pl. 10, fi gs 1–6.<br />

R e m a r k s. These pollen grains represent<br />

I type <strong>of</strong> sacci attachment to <strong>the</strong> corpus (after<br />

Sivak 1976).<br />

(47) Cathayapollis potoniei (Sivak 1976)<br />

Ziembińska-Tworzydło in Stuchlik et al. 2002<br />

Pl. 5, fi g. 2a, b.<br />

1953 Pityosporites microalatus (Potonié) n. comb.,<br />

Thomson & Pfl ug, p. 67, pl. 5, fi gs 51, 53, 59.<br />

1954 Pinus haploxylon classical type Rudolph; Doktorowicz-Hrebnicka,<br />

p. 64.<br />

1976 Cathaya potoniei sp. nov., Sivak, p. 272, pl. 13,<br />

fi gs 1–10.<br />

2002 Cathayapollis potoniei (Sivak) Ziembińska-<br />

Tworzydło comb. nov.; Stuchlik et al., p. 17,<br />

pl. 15, fi gs 1–10.<br />

R e m a r k s. These pollen grains represent III<br />

type <strong>of</strong> sacci attachment to <strong>the</strong> corpus (after<br />

Sivak 1976).<br />

(48) Cathayapollis wilsonii (Sivak 1976)<br />

Ziembińska-Tworzydło in Stuchlik et al. 2002<br />

Pl. 5, fi g. 3a, b<br />

1960 Pinus type Haploxylon Rudolph = Pollenites<br />

microalatus, Oszast, p. 15, pl. 5, fi gs 3, 4.<br />

1971 Pityosporites alatus (Potonié) Thomson & Pfl ug<br />

[=Pinus alata (Potonié) n. comb.]; Krutzsch,<br />

p. 51, pl. 2, fi gs 2, 3, 8, 9.<br />

1976 Cathaya wilsonii sp. nov., Sivak, p. 270, pl. 12,<br />

fi gs 1–10.

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