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Middle Miocene palynoflora of the Legnica lignite deposit complex ...

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1953 Verrucatosporites favus (Potonié) n. comb.,<br />

Thomson & Pfl ug, p. 60, pl. 3, fi gs 52–55, pl. 4,<br />

fi gs 1–3.<br />

1959 Reticuloidosporis (Polypodiisporites) favus<br />

(Potonié) n. comb., Krutzsch, p. 215, pl. 42, fi gs<br />

467–470.<br />

R e m a r k s. Only one spore <strong>of</strong> this taxon was<br />

found in <strong>the</strong> studied material in <strong>the</strong> Lusatian<br />

seam. Morphologically, it is similar to spores<br />

<strong>of</strong> <strong>the</strong> recent Dennstaedtiaceae, especially to<br />

genus Paesia. This taxon represents cosmopolitan<br />

(P/A) climatic element, and occurs<br />

in Europe in <strong>the</strong> <strong>Middle</strong> Eocene to <strong>Miocene</strong><br />

fossil palyn<strong>of</strong>l oras. It was also reported from<br />

<strong>the</strong> Lower and <strong>Middle</strong> <strong>Miocene</strong> <strong>of</strong> central and<br />

western Poland.<br />

Dennstaedtiaceae (16 genera) occur worldwide<br />

in tropical to temperate areas. Paesia<br />

is distributed in tropical South and <strong>Middle</strong><br />

America, Malaysia and Pacifi c Islands (Stuchlik<br />

et al. 2001).<br />

Perinomonoletes Krutzsch 1967<br />

(20) Perinomonoletes pliocaenicus<br />

Krutzsch 1967<br />

Pl. 2, fi g. 8<br />

1967 Perinomonoletes pliocaenicus n. sp., Krutzsch,<br />

p. 222, pl. 87, fi gs 2–9.<br />

Spores monolete, bilateral, 45–50 × 25–30<br />

μm in size. Laesura poorly visible. Exine 1.0–<br />

1.5 μm thick. Perine smooth, waved, protruding<br />

up to 3.5 μm from exine surface.<br />

(21) Perinomonoletes cf. goersbachensis<br />

Krutzsch 1967<br />

Pl. 2, fi g. 11<br />

1967 Perinomonoletes goersbachensis n. sp., Krutzsch,<br />

p. 222, pl. 87, fi gs 10–12.<br />

R e m a r k s. Several spores resembling in<br />

structure Perinomonoletes pliocaenicus, but<br />

more than 55 μm long, nearest <strong>the</strong> fossil taxon<br />

P. goersbachensis were found.<br />

(22) Perinomonoletes spicatus Nagy 1973<br />

Pl. 2, fi g. 9a, b<br />

1973 Perinomonoletes spicatus n. sp., Nagy, p. 454,<br />

pl. 2, fi gs 6, 7.<br />

Spores similar in structure to Perinomonoletes<br />

pliocaenicus, 32–55 μm long. Perine with<br />

delicate “spines”.<br />

(23) Perinomonoletes sp. 1<br />

Pl. 2, fi g. 10<br />

19<br />

Spores similar to <strong>the</strong> above-mentioned ones,<br />

but perine thin, very delicate, protruding up to<br />

a dozen or so μm from exine surface.<br />

R e m a r k s. Spores <strong>of</strong> <strong>the</strong> morphological genus<br />

Perinomonoletes approach those <strong>of</strong> recent<br />

members <strong>of</strong> Polypodiaceae s.l.; particularly<br />

Aspleniaceae, Blechnaceae and Aspidiaceae<br />

(Nayar & Devi 1964a–c, Kremp & Kawasaki<br />

1972), and <strong>the</strong> genera Asplenium L., Athyrium<br />

Roth, Bolbitis Schott, Byrsopteris Morton,<br />

Cornopteris Nakai, Ctenitis C. Chr., Cyclosorum<br />

Link, Diplazium Sw., Dryopteris Adans.,<br />

Egenolfi a Schott, Lastrea Bory, Lithostegia<br />

Ching, Peranema Don., and Polystichum Roth<br />

( Krutzsch 1967, Thiele-Pfeiffer 1980). They<br />

were reported from <strong>the</strong> Eocene and Pliocene<br />

(Krutzsch 1967), as well <strong>the</strong> <strong>Miocene</strong> <strong>deposit</strong>s<br />

<strong>of</strong> Germany (Thiele-Pfeiffer 1980) and Hungary<br />

(Nagy 1985). In <strong>the</strong> analysed material<br />

<strong>the</strong>y occurred only in two samples from <strong>the</strong><br />

Komorniki pr<strong>of</strong>i le.<br />

Classis PTEROPSIDA<br />

incerte sedis<br />

Toroisporis Krutzsch 1959<br />

(24) Toroisporis (Toroisporis) teupitzensis<br />

medioris Krutzsch 1962<br />

Pl. 3, fi g. 1a, b<br />

1962a Toroisporis (Toroisporis) teupitzensis medioris<br />

n. subfsp., Krutzsch, p. 80, pl. 33, fi gs 1–14.<br />

R e m a r k s. Spores <strong>of</strong> this taxon are slightly<br />

similar to those <strong>of</strong> <strong>the</strong> recent families Cya<strong>the</strong>aceae,<br />

Dipteridaceae, Pteridaceae and Lygodiaceae.<br />

They represent palaeotropical/warmtemperate<br />

element (P/A1), and occur in Europe<br />

in <strong>the</strong> Lower – <strong>Middle</strong> <strong>Miocene</strong>. They were also<br />

found in <strong>the</strong> Upper Oligocene to Lower Pliocene<br />

<strong>deposit</strong>s in south-western Poland (Stuchlik<br />

et al. 2001). In <strong>the</strong> studied material only<br />

one spore was found in <strong>the</strong> Mużaków series.<br />

(25) Toroisporis (Toroisporis)?<br />

pliocaenicus (Thiergart 1940)<br />

Krutzsch 1962<br />

Pl. 3, fi g. 2<br />

1940 Sporites neddeni f. pliocaenicus Thiergart, p. 25,<br />

pl. 1, fi g. 2.

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