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Middle Miocene palynoflora of the Legnica lignite deposit complex ...

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1959 Baculatisporites major (Raatz) n. comb., Krutzsch,<br />

p. 140.<br />

1967 Baculatisporites primarius major Raatz; Krutzsch,<br />

p. 56, pl. 10, fi gs 1–6.<br />

1994b Osmundacidites primarius major (Raatz) Ziembińska-Tworzydło<br />

comb. nov.; Ziembińska-<br />

Tworzydło et al., p. 12, pl. 4, fi g. 10a, b.<br />

2001 Baculatisporites major (Raatz) Krutzsch; Stuchlik<br />

et al., p. 30, pl. 24, fi gs 1–3.<br />

R e m a r k s. According to Krutzsch (1967)<br />

spores <strong>of</strong> this taxon are similar to <strong>the</strong> recent<br />

Osmunda vachelli Hook., and O. presliana<br />

J. Sm. They occur in Europe in <strong>the</strong> Upper Oligocene<br />

to Pliocene <strong>deposit</strong>s, and are common<br />

in <strong>the</strong> Polish <strong>Middle</strong> <strong>Miocene</strong>. They are classifi<br />

ed to cosmopolitan (P/A) climatic element<br />

(Stuchlik et al. 2001). In <strong>the</strong> studied material<br />

<strong>the</strong>se spores were <strong>of</strong>ten found.<br />

(12) Baculatisporites primarius<br />

(Wolff 1934) Pfl ug & Thomson<br />

in Thomson & Pfl ug 1953<br />

Pl. 1, fi g. 14a, b<br />

1934 Sporites primarius n. sp., Wolff, p. 66, pl. 5,<br />

fi g. 8.<br />

1953 Baculatisporites primarius (Wolff) Pfl ug & Thomson<br />

n. comb.; Thomson & Pfl ug, p. 56, pl. 2,<br />

fi g. 51.<br />

2001 Baculatisporites primarius (Wolff) Pfl ug & Thomson<br />

in Thomson & Pfl ug; Stuchlik et al., p. 31,<br />

pl. 25, fi g. 11, pl. 26, fi gs 1–8, pl. 27, fi gs 1–7.<br />

R e m a r k s. Krutzsch (1967) distinguished<br />

four subspecies <strong>of</strong> Baculatisporites primarius<br />

(crassiprimarius, oligocenicus, primarius, and<br />

semiprimarius) and compared <strong>the</strong>m e.g. with<br />

<strong>the</strong> recent species Osmunda banksiaefolia<br />

(Pr.) Kuhn. (from East Asia), O. interrupta<br />

Michx. (from North America), O. javanica Bl.,<br />

O. bromeliifolia (Pr.) Copel. (from Ceylon),<br />

and Osmunda vachelli Hook. Stuchlik and coauthors<br />

(2001) included <strong>the</strong>se subspecies into<br />

a synonym list. Baculatisporites primarius<br />

spores occur in Europe in <strong>the</strong> Upper Oligocene<br />

to Pliocene <strong>deposit</strong>s, and are common in <strong>the</strong><br />

Polish Neogene. They represent cosmopolitan<br />

(P/A) climatic element (Stuchlik et al. 2001). In<br />

<strong>the</strong> studied material <strong>the</strong>se spores were found<br />

regularly.<br />

In addition, some spores <strong>of</strong> fossil species<br />

Baculatisporites nanus (Wolff 1934)<br />

Krutzsch 1959 were found in <strong>the</strong> Komorniki<br />

pr<strong>of</strong>i le. This taxon represents cosmopolitan<br />

(P/A) climatic element, and occurs in <strong>the</strong><br />

Upper Oligocene to Pliocene fossil fl oras. It is<br />

17<br />

common all over Poland in <strong>the</strong> <strong>Middle</strong> <strong>Miocene</strong><br />

<strong>deposit</strong>s (Stuchlik et al. 2001).<br />

Rugulatisporites Pfl ug & Thomson 1953<br />

(13) Rugulatisporites quintus Pfl ug<br />

& Thomson in Thomson & Pfl ug 1953<br />

Pl. 1, fi g. 13<br />

1953 Rugulatisporites quintus Pfl ug & Thomson n.<br />

sp.; Thomson & Pfl ug, p. 56, pl. 2, fi g. 46.<br />

1967 Baculatisporites quintus (Thomson & Pfl ug) n.<br />

comb., Krutzsch, p. 48, pl. 6–8.<br />

1985 Osmundacidites quintus (Pfl ug & Thomson)<br />

n. comb. ssp. Quintus, Nagy, p. 75, pl. 13, fi gs<br />

1, 2.<br />

2001 Rugulatisporites quintus Pfl ug & Thomson in<br />

Thomson & Pfl ug; Stuchlik et al., p. 49, pl. 28,<br />

fi gs 1–5, pl. 29, fi gs 1–4.<br />

Remarks. Rugulatisporites quintus represents<br />

subtropical/warm-temperate element<br />

(P2/A1). In Europe it occurs in <strong>the</strong> Upper Oligocene<br />

to Pliocene <strong>deposit</strong>s. It is common in <strong>the</strong><br />

Polish <strong>Miocene</strong> and Pliocene. Morphologically,<br />

<strong>the</strong>se fossil spores are most similar to those <strong>of</strong><br />

<strong>the</strong> recent Osmunda regalis L. ( Krutzsch 1967,<br />

Stuchlik et al. 2001). The fossil spores <strong>of</strong> Rugulatisporites<br />

quintus were <strong>of</strong>ten encountered<br />

in <strong>the</strong> analysed material, mainly in samlpes<br />

above <strong>the</strong> Lusatian seam.<br />

Nowadays <strong>the</strong> genus Osmunda comprises<br />

14 species occurring in humid forests in temperate<br />

zone through swampy tropical regions<br />

<strong>of</strong> <strong>the</strong> nor<strong>the</strong>rn hemisphere. O. regalis is distributed<br />

in subtropical and temperate regions<br />

<strong>of</strong> both hemispheres.<br />

Familiae ?SCHIZEACEAE,<br />

?CYATHEACEAE<br />

Monoleiotriletes Krutzsch 1959<br />

(14) Monoleiotriletes gracilis<br />

Krutzsch 1959<br />

Pl. 2, fi g. 2a, b<br />

1959 Monoleiotriletes gracilis n. sp., Krutzsch, p. 65,<br />

pl. 4, fi g. 24.<br />

2001 Monoleiotriletes gracilis Krutzsch; Stuchlik<br />

et al., p. 43, pl. 22, fi g. 6a, b.<br />

R e m a r k s. Only one spore <strong>of</strong> this taxon was<br />

found in <strong>the</strong> Komorniki pr<strong>of</strong>i le. This taxon<br />

occurs in Europe in <strong>the</strong> <strong>Middle</strong> Eocene to<br />

<strong>Middle</strong> <strong>Miocene</strong> <strong>deposit</strong>s. It is reported from<br />

<strong>the</strong> Lower <strong>Miocene</strong> <strong>of</strong> south-western Poland.

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