supporting lamella
supporting lamella
supporting lamella
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HISTOLOGY AND DEVELOPMENT OF MYRIOTHELA PHRYGTA. 513<br />
endoderm are undistinguishable. This grows while at the<br />
same time its elements lose their distinctness and become<br />
highly charged with spherical masses of stored nutriment,<br />
resembling in many particulars the nutritive spheres of the<br />
general endoderm. As it grows it pushes the perisarc before<br />
it, and ultimately forms a rounded egg-like mass attached to<br />
the parent body by a short thick pedicle (fig. 13). From<br />
this the young Myriothela is developed (fig. 13). All connection<br />
with the body of the parent is lost at a very early<br />
period, almost before the bud has re-formed its ectoderm and<br />
endoderm and enteric cavity. It remains attached to the<br />
perisarc, however, by a sucker-like arrangement at the aboral<br />
pole until it is fully formed.<br />
As will be seen, the formation of a gonophore is, in its<br />
earliest stages, essentially similar to this method of budding.<br />
In other words, the gonophore is a true bud which, like the<br />
other buds, is derived from a blastema formed by a fusion of<br />
ectodermal and endodermal elements. The difference, however,<br />
lies in the fact that in the case of the gonophore bud, after it<br />
is a well-formed structure, a group of the primitive germ-cells<br />
make their way into it.<br />
The first stage in the growth of a gonophore is shown<br />
in fig. 8. The ectoderm of the gonophore-bearing region<br />
becomes thickened over a small surface, the increase in<br />
thickness being due largely to an accumulation of the<br />
primitive germ-cells, but partly to an increase in the cells<br />
carrying hyaline masses. At the same time nematocysts disappear<br />
in that region of the ectoderm, though they may occur<br />
in their usual profusion in close proximity. In the next stage<br />
the basement membrane is absorbed or ruptured (figs. 7 and<br />
9), I cannot determine which, and a tongue of endodermcells<br />
pushes its way into the ectoderm and through the deepest<br />
layer. Thus the cluster of primitive germ-cells come to lie<br />
not on its apex, but, generally, asymmetrically disposed on<br />
one side.<br />
The removal of the <strong>supporting</strong> <strong>lamella</strong> is, I am inclined to<br />
think, mainly a process of solution, since scattered rounded<br />
VOL. XXXII, PART IV.—NEW SEE. M M