06.04.2013 Views

supporting lamella

supporting lamella

supporting lamella

SHOW MORE
SHOW LESS

Create successful ePaper yourself

Turn your PDF publications into a flip-book with our unique Google optimized e-Paper software.

528 W. B. HARDY.<br />

ingested by the apical cells, both in the body of the animal and<br />

in the proximal region of the blastostyles.<br />

Towards the close of digestion a few free cells appear in<br />

the somatic fluid. Each is rounded and composed of darkstaining<br />

protoplasm embedding a nucleus with contained<br />

nucleolus.<br />

Before turning to the further fate of the food, that is to say<br />

before considering the process of absorption, it will be well to<br />

describe more particularly the contents and characters of the<br />

vacuolate cells.<br />

These when unloaded with nutritive spheres present the<br />

appearance shown in fig. 18, where the cell is seen to consist<br />

of a thin pellicle of protoplasm surrounding a large central<br />

vacuole and embedding a nucleus. The protoplasm stains only<br />

slightly. The outline of the cell is exceedingly sharply defined<br />

by some staining material which has almost the appearance of a<br />

cuticle. The process of loading with nutritive spheres is remarkable,<br />

and essentially similar to that described by Miss Greenwood<br />

as occurring in Hydra. The protoplasm at one point<br />

develops a small vacuole which increases in size, and bulges into<br />

the large vacuole. In this the nutritive sphere is formed from<br />

the turbid semi-fluid material which first fills it. This process<br />

continues until the whole of the cell becomes occupied by small<br />

vacuoles, each containing a nutritive sphere. The size of the<br />

cell, therefore, does not necessarily vary according to the amount<br />

of reserve nutriment it contains. This, however, only holds<br />

good for the vacuolate cells of the body. In the blastostyles<br />

they are slightly different. In the first place the large central<br />

vacuole is not developed, with the consequence that when the<br />

cell discharges its nutritive spheres it frequently shrinks to the<br />

condition of a cell with dense, non-vacuolated protoplasm, which<br />

stains deeply with picro-carmine. In fig. 22, at 1 is a cell<br />

without nutritive spheres, and at 2 one partly filled with<br />

vacuoles containing nutritive spheres.<br />

But the discharge of the nutritive spheres does not always<br />

leave the cells smaller and solid. The vacuoles may persist<br />

(fig. 22, 4), and the result is a remarkable cell with bubbly

Hooray! Your file is uploaded and ready to be published.

Saved successfully!

Ooh no, something went wrong!