A TAXONOMIC REVISION OF HYMENOPHYLLACEAE

A. Ebihara et al.: A taxonomic revision of Hymenophyllaceae
densely hairy with long bristle-like light reddish hairs. Blades tripinnate to quadripinnate,
elliptic to narrowly ovate, 40 by 16 cm, venation anadromous, false veinlets
absent, internal cell walls thin and straight. Sori paratactic, campanulate, lips truncate,
or sometimes bilabiate, receptacles long-exserted.
Distribution — The Paleotropics (Asia to the Pacific); c. 5 species.
Habitat — Usually epilithic on moist rocks, occasionally epiphytic on tree trunks.
Chromosome base number — x = 36.
Note — Corresponding to the Ca clade in Ebihara et al. (submitted).
RepResentative species:
New combination:
Callistopteris superba (Backh. ex T. Moore) Ebihara & K. Iwats., comb. nov. [based on Trichomanes
superbum Backh. ex T. Moore (1862) 44].
Other species:
Callistopteris apiifolia (C. Presl) Copel.; C. baldwinii (D.C. Eaton) Copel.; C. baueriana (Endl.)
Copel.; C. polyantha (Hook.) Copel.
ACKNOWLEDGEMENTS
We are grateful to Ms. E. Wood (Harvard University Herbaria) for checking and correcting the
English manuscript; to Dr. P.D. Bostock (Queensland Herbarium) for checking the Latin diagnosis;
to the directors and curators of herbaria (BM, K, L and P) for supporting our investigation of their
specimens; to Dr. N. Murakami and Dr. H. Nishida for providing some literature; to Dr. Leon Perrie
(Museum of New Zealand Te Papa Tongarewa) for giving advices about the replacement for Cardiomanes
reniforme and to the anonymous reviewer for valuable comments on the manuscript. This
study was partly supported by the JSPS Grant for Oversea Research No. 12575012 (for K.I.) and
by JSPS Fellows (for A.E.).
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