A TAXONOMIC REVISION OF HYMENOPHYLLACEAE

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BLUMEA — Vol. 51, No. 2, 2006
Habitat — Epilithic or low-epiphytic.
Note — Corresponding to the Le subclade in Ebihara et al. (submitted).
RepResentative species:
New combinations:
Didymoglossum hildebrandtii (Kuhn) Ebihara & Dubuisson, comb. nov. [based on Trichomanes
hildebrandtii Kuhn (1879) 70]; D. lorencei (Tardieu) Ebihara & Dubuisson, comb. nov. [based on
Trichomanes lorencei Tardieu (1977) 147]; D. motleyi (Bosch) Ebihara & K. Iwats., comb. nov. [based
on Trichomanes motleyi Bosch (1861b) 145]; D. sublimbatum (Müll.Berol.) Ebihara & K. Iwats.,
comb. nov. [based on Trichomanes sublimbatum Müll.Berol. (1854) 737]; D. tahitense (Nadeaud)
Ebihara & K. Iwats., comb. nov. [based on Trichomanes tahitense Nadeaud (1873) 18].
Other species:
Didymoglossum angustifrons Fée; D. curtii (Rosenst.) Pic.Serm.; D. exiguum (Bedd.) Copel.;
D. gourlianum (Grev. ex J. Sm.) Pic.Serm.; D. hymenoides (Hedw.) Copel.; D. krausii (Hook. &
Grev.) C. Presl; D. liberiense (Copel.) Copel.; D. lineolatum Bosch; D. melanopus (Baker) Copel.;
D. membranaceum (L.) Vareschi; D. nummularium Bosch; D. ovale E. Fourn.; D. petersii (A. Gray)
Copel.; D. pinnatinervium (Jenman) Pic.Serm.; D. punctatum (Poir.) Desv.; D. pusillum (Sw.) Desv.;
D. reptans (Sw.) C. Presl; D. rhipidophyllum (Sloss.) Pic.Serm.
2. Subgenus Microgonium (C. Presl) Ebihara & K. Iwats., comb. nov.
Based on Microgonium C. Presl (1843) 19, t. VI, f. A, B. — Hemiphlebium C. Presl sect. Microgonium
(C. Presl) Prantl (1875) 46. — Trichomanes L. sect. Microgonium (C. Presl) H. Christ (1897) 24.
— Lectotype: Microgonium cuspidatum (Willd.) C. Presl (= Didymoglossum cuspidatum (Willd.)
Ebihara & Dubuisson, comb. nov. [based on Trichomanes cuspidatum Willd. (1810) 499]) (selected
by Christensen (1906) XIV).
Rhizomes long-creeping, frequently branching, filiform, to 0.5 mm diam., densely covered
with dark hairs, protostele collateral to regressed (see Ebihara et al., submitted),
cortex petiole-like, roots absent, root-like shoots present. Stipes often quite reduced,
to 1 cm, at a distance from the adjacent ones. Blades usually simple, or sometimes
lobed to pinnatifid, elliptic to narrowly ovate, 3 by 2 cm, venation catadromous, continuous
submarginal false veinlets present, longitudinal false veinlets parallel to the
true veins present, internal cell walls thin and straight, margins entire and glabrous.
Sori epitactic or pantotactic, often immersed in the laminae, tubular, lips often dilate,
receptacles exserted.
Distribution — Throughout the tropics, mainly in the Old World; more than 10 species.
Habitat — Epilithic or low-epiphytic.
Note — Corresponding to the Mg subclade in Ebihara et al. (submitted).
RepResentative species:
New combinations:
Didymoglossum bimarginatum (Bosch) Ebihara & K. Iwats., comb. nov. [based on Microgonium
bimarginatum Bosch (1861b) 143]; D. ekmanii (Wess. Boer) Ebihara & Dubuisson, comb. nov.
[based on Trichomanes ekmanii Wess.Boer (1962) 319, f. 33a, b]; D. godmanii (Hook.) Ebihara &
Dubuisson, comb. nov. [based on Trichomanes godmanii Hook. in Baker (1866) 337, t. 8, f. A]; D. kap-
plerianum (Sturm) Ebihara & Dubuisson, comb. nov. [based on Trichomanes kapplerianum Sturm in
Martius (1859) 276]; D. kirkii (Hook.) Ebihara & Dubuisson, comb. nov. [based on Trichomanes kirkii
Hook. in Hook. & Baker (1867) 78]; D. lenormandii (Bosch) Ebihara & Dubuisson, comb. nov. [based
on Trichomanes lenormandii Bosch (1861b) 144]; D. mindorense (H. Christ) K. Iwats., comb. nov.
[based on Trichomanes mindorense H. Christ (1908a) 270]; D. pygmaeum (C. Chr.) Ebihara & Dubuisson,
comb. nov. [based on Trichomanes pygmaeum C. Chr. (1920) 10, f. 2]; D. sinuatum (Bonap.)