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68<br />

Balearic Islands<br />

EUROPE<br />

SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY<br />

FIGURE 1.—Map of the Mediterranean region showing the position and nomenclature of the Balearic Islands<br />

used in this paper. Eivissa is the Catalan name for the island known in other languages as Ibiza.<br />

native, prehuman avifaunas as those of the other islands, we<br />

have excluded Menorca and Formentera from our analysis. The<br />

numbers of fossil bones presented above, together with data on<br />

species distribution in the different deposits, allow a rough estimate<br />

of relative species abundance on the islands, taphonomic<br />

biases included.<br />

The very different vertebrate paleofaunas of the Mediterranean-basin<br />

archipelagos are of interest for comparison with<br />

other insular faunas worldwide. Herein we present a preliminary<br />

analysis of the paleoecology of the late Pleistocene bird<br />

communities of the western Mediterranean islands based on<br />

the known trophic ecology of the different species recorded on<br />

the islands. We emphasize parallels with other known prehuman<br />

native-bird communities worldwide. The Pleistocene bird<br />

communities of the western Mediterranean mainly include extant<br />

species, even though many have recently vanished from<br />

several or all the islands considered. They also include at least<br />

four or five extinct species: Tyto balearica Mourer-Chauvire,<br />

Alcover, Moya, and Pons, a nonendemic species found on<br />

Mallorca, Menorca, Corsica (Mourer-Chauvire, pers. comm.,<br />

1996), and in some mainland deposits from France and the<br />

Iberian Peninsula; Bubo insularis Mourer-Chauvire and Weesie,<br />

endemic to Corsica and Sardinia; an undescribed species of<br />

Athene endemic to Corsica (Mourer-Chauvire, pers. comm.,<br />

1996); an undescribed species of Rallus endemic to Eivissa;<br />

and Grus primigenia Milne-Edwards, a nonendemic species<br />

widely distributed during the European Pleistocene, also<br />

present on Eivissa and Mallorca, which is probably conspecific<br />

with Grus grus (J.R. Stewart, pers. comm., 1996). Some<br />

taxa previously considered as possible endemics (e.g., Fringillidae,<br />

species undescribed, and Corvidae, species undescribed;<br />

see Alcover et al., 1992) have been deleted after taxonomic<br />

reappraisal. Nomenclature for binomials of modern<br />

birds follows Sibley and Monroe (1990).<br />

Even though the determination of the feeding habits and specializations<br />

of extinct species may be imprecise or speculative,<br />

the allocation of these species to broad trophic categories can<br />

allow us to undertake a more general analysis. These kinds of<br />

inferences are applicable to all the extinct Mediterranean species<br />

considered above, but they cannot be extended indiscriminately,<br />

especially in the case of bizarre small Passeriformes<br />

without known analogs (e.g., Vangulifer from Maui, Hawaiian<br />

Islands; James and Olson, 1991).<br />

ACKNOWLEDGMENTS.—We are indebted to Storrs Olson,<br />

Helen James, Dave Steadman, and Anna Traveset for their<br />

comments, ideas, and useful discussions. Damia Jaume and

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