PDF (Lo-Res) - Smithsonian Institution Libraries
PDF (Lo-Res) - Smithsonian Institution Libraries PDF (Lo-Res) - Smithsonian Institution Libraries
46 SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY P. urinatrix exsul (Common Diving-petrel): MNZ 908S, 17623-17625, 18096 (Auckland Id.); MNZ 24785, 24786 (Heard Id.). P urinatrix urinatrix: MNZ 546S, 547S, 13610, 13613, 13614, 16601, 17198, 17202 (Wellington-area beaches, New Zealand). P. georgicus (South Georgian Diving-petrel): MNZ 24790, 24816-24822 (Heard Id.). Catharacta skua lonnbergi (Subantarctic (Brown or Southem Great) Skua): MNZ 13568, 18332 (Auckland Id.); MNZ 18900 (Southeast Id., Chatham Ids.); MNZ 23702 (Campbell Id.); MNZ 24698, 24699 (Snares Id.). C. maccormicki (South Polar or Antarctic Skua): MNZ 13320 (Franklin Id.); MNZ 22955, 22956 (Cape Evans); MNZ 22935, 22947, 23035, 23158 (Scott Base). Systematics and Comparative Osteology Table 1 lists the species determined in the Martinez collection, the number of bones identified as belonging to each species, and the minimum number of individuals (MNI) per species. The fossils described herein provide evidence for the previous fauna having at least 20 species of seabird and one duck. To this total can be added the records of a presumably vagrant rail, Crex crex, and a domestic fowl, Gallus gallus (Linnaeus), the latter found in a site used by sailors for shelter. Eudyptes chrysocome All penguin bones are referred to Eudyptes. They are the same size as bones of the Fiordland Penguin {E. pachyrhynchus), Snares Penguin {E. robustus), and Eastern Rockhopper Penguin {E. chrysocome filholi). Bones of the Macaroni Penguin {E. chrysolophus) and the Erect-crested Penguin {E. sclateri) are much larger. Because the Northern Rockhopper Penguin {E. chrysocome moseleyi) breeds on Amsterdam in large numbers (Jouventin, 1994), the bones are referred to that species. Diomedea chlororhynchos Most albatross bones were removed from the collection before THW examined them. A mandible from site 26, and eight bones of one individual from site 2.1, however, are all indistinguishable from the Yellow-nosed Albatross {Diomedea chlororhynchos) and are referred to that species. Large numbers of these albatrosses currently breed on Amsterdam (Jouventin et al., 1984). Diomedea amsterdamensis The Martinez collection contained numerous remains of Diomedea amsterdamensis that were removed prior to THW's examination of the collection. These were reported on by Jouventin et al. (1989), who recorded bones representing 81 indi- TABLE 1.—List of species, number of bones, and minimum number of individuals (MNI) calculated as the sum of MNI in individual sites, as defined by the collectors, represented in the fossils from Amsterdam Island. Species Eudyptes chrysocome Diomedea sp. Diomedea amsterdamensis Diomedea chlororhynchos Procellaria cinerea Pterodroma macroptera Pterodroma mollis Pterodroma baraui Pterodroma baraui/arminjoniana Pterodroma arminjoniana Pterodroma sp. ?aterrima Pachyptila macgillivrayi Pachyptila cf. turtur Pachyptila cf. desolata Pachyptila sp. chicks Pelecanoides urinatrix Puffinus assimilis Puffinus carneipes Puffinus cf. griseus Pelagodroma marina Fregetta grallaria Oceanites sp. Hydrobatidae sp. indet. Catharacta skua hamiltoni Anas marecula Gallus gallus Total Number of bones 281 42 80 9 980 3711 4766 263 11 6 2 9235 57 1 7 106 1882 21 2 1780 72 17 5 179 44 3 23562 MNI 21 3 9(8 ir 2 117 282 375 31 3 4 2 731 12 1 2 29 152 1 1 206 25 4 2 19 24(33)* 1 2059 "Jouventin et al. (1989) recorded 81 individuals of D. amsterdamensis, and Olson and Jouventin (1996) reported a minimum of 33 individuals of Anas marecula, and there is probably overlap between those samples and what we examined in the study. viduals of D. amsterdamensis from several unspecified sites distributed between 80 m and 700 m a.s.l., including 52 individuals from a single site at 332 m, which is probably site 4. About 80 bones of a large species of albatross remained in the collection, however, and are referred to D. amsterdamensis because they do not match any other species. Diomedea amsterdamensis is restricted to breeding sites above 400 m (Jouventin etal., 1989). Pterodroma FIGURES 5-8 A preliminary examination of the Pterodroma bones in the collection revealed the predominance of two different species that differed mainly in size. In order to characterize these better, a large sample was measured, revealing that both species have apparently normal distributions for all bones (Figures 7, 8; Appendices 1,2). Species in the size range of the larger taxon include the Great-winged or Grey-faced Petrel {Pterodroma macroptera), Juan Fernandez Petrel {P. externa), Providence Petrel {P. solandri), Magenta Petrel {P. magentae), and White-necked
NUMBER 89 FIGURE 5.—Photographs of Pterodroma mollis skulls in dorsal (left) and lateral (right) views. A,D, modem specimen MNZ 22424, Gough Island; B,C, fossil specimen from site 18.7. Scale bar=5 cm. FIGURE 6.—Photographs of Pterodroma macroptera skulls in dorsal (left) and lateral (right) views. A,D, modern specimen CSIRO PROS237, Eclipse Island, Western Australia; B,c, specimen from fossil site 18.3. Scale bar=5 cm. Petrel {P. cervicalis). The smaller taxon is about the size of a Soft-plumaged Petrel {P. mollis), Mottled Petrel {P. inexpectata), Black-winged Petrel {P. nigripennis), or Kerguelen Petrel (P. brevirostris). Bones of the Atlantic Petrel {P. incerta), White-headed Petrel {P. lessoni), and Tahiti Petrel {P. rostrata) are larger than those of any Pterodroma from Amsterdam, and 47
- Page 6 and 7: ABSTRACT Olson, Storrs L., editor.
- Page 8 and 9: EARLY WATERFOWL (ANSERIFORMES) AND
- Page 10 and 11: v m SMITHSONIAN CONTRIBUTIONS TO PA
- Page 12 and 13: localities, all of them situated in
- Page 14 and 15: Bone of Sus scrofa Linnaeus, introd
- Page 16 and 17: duboisi are larger than the largest
- Page 18 and 19: 8 iver (1897) but afterward were tr
- Page 20 and 21: 10 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 22 and 23: 12 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 24 and 25: 14 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 26 and 27: 16 Cor. Hum. Uln. Cpm. Fern. Tbt. T
- Page 28 and 29: 18 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 30 and 31: 20 sometatarsus are proportionally
- Page 32 and 33: 22 Cor. Hum. Uln. Cpm. Fern. Tbt. T
- Page 34 and 35: 24 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 36 and 37: 26 Cor. Hum. Cpm. Fern. Tbt. Tmt. P
- Page 38 and 39: 28 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 40 and 41: 30 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 42 and 43: 32 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 44 and 45: 34 ability in such a way that it ca
- Page 46 and 47: 36 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 48 and 49: 38 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 50 and 51: 40 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 52 and 53: 42 2km SMITHSONIAN CONTRIBUTIONS TO
- Page 54 and 55: Site 13 Research Base ENTRANCE Lowe
- Page 58 and 59: 48 the last has unusually slender w
- Page 60 and 61: 50 Si 5 15i 10- 5- 20 J 15 I 10 f 5
- Page 62 and 63: 52 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 64 and 65: 54 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 66 and 67: 56 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 68 and 69: 58 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 70 and 71: 60 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 72 and 73: 62 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 74 and 75: 64 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 77 and 78: Comparison of Paleoecological Patte
- Page 79 and 80: NUMBER 89 69 TABLE 1.—Vertebrate
- Page 81 and 82: NUMBER 89 71 Mallorca, probably in
- Page 83: NUMBER 89 Alcover, J.A. 1989. Les a
- Page 86 and 87: 76 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 88 and 89: 78 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 90 and 91: 80 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 92 and 93: 82 SMITHSONIAN CONTRIBUTIONS TO PAL
- Page 95 and 96: The History of the Chatham Islands'
- Page 97 and 98: NUMBER 89 87 Species Common Name Pe
- Page 99 and 100: NUMBER 89 89 NZA 797,1930,1934 Unco
- Page 101 and 102: NUMBER 89 91 anoramphus spp.), Chat
- Page 103 and 104: NUMBER 89 93 TABLE 2.—Land snails
- Page 105 and 106: NUMBER 89 95 counterparts, with som
46 SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY<br />
P. urinatrix exsul (Common Diving-petrel): MNZ 908S,<br />
17623-17625, 18096 (Auckland Id.); MNZ 24785, 24786<br />
(Heard Id.).<br />
P urinatrix urinatrix: MNZ 546S, 547S, 13610, 13613,<br />
13614, 16601, 17198, 17202 (Wellington-area beaches, New<br />
Zealand).<br />
P. georgicus (South Georgian Diving-petrel): MNZ 24790,<br />
24816-24822 (Heard Id.).<br />
Catharacta skua lonnbergi (Subantarctic (Brown or Southem<br />
Great) Skua): MNZ 13568, 18332 (Auckland Id.); MNZ<br />
18900 (Southeast Id., Chatham Ids.); MNZ 23702 (Campbell<br />
Id.); MNZ 24698, 24699 (Snares Id.).<br />
C. maccormicki (South Polar or Antarctic Skua): MNZ<br />
13320 (Franklin Id.); MNZ 22955, 22956 (Cape Evans); MNZ<br />
22935, 22947, 23035, 23158 (Scott Base).<br />
Systematics and Comparative Osteology<br />
Table 1 lists the species determined in the Martinez collection,<br />
the number of bones identified as belonging to each species,<br />
and the minimum number of individuals (MNI) per species.<br />
The fossils described herein provide evidence for the<br />
previous fauna having at least 20 species of seabird and one<br />
duck. To this total can be added the records of a presumably<br />
vagrant rail, Crex crex, and a domestic fowl, Gallus gallus<br />
(Linnaeus), the latter found in a site used by sailors for shelter.<br />
Eudyptes chrysocome<br />
All penguin bones are referred to Eudyptes. They are the<br />
same size as bones of the Fiordland Penguin {E. pachyrhynchus),<br />
Snares Penguin {E. robustus), and Eastern Rockhopper<br />
Penguin {E. chrysocome filholi). Bones of the Macaroni Penguin<br />
{E. chrysolophus) and the Erect-crested Penguin {E. sclateri)<br />
are much larger. Because the Northern Rockhopper Penguin<br />
{E. chrysocome moseleyi) breeds on Amsterdam in large<br />
numbers (Jouventin, 1994), the bones are referred to that species.<br />
Diomedea chlororhynchos<br />
Most albatross bones were removed from the collection before<br />
THW examined them. A mandible from site 26, and eight<br />
bones of one individual from site 2.1, however, are all indistinguishable<br />
from the Yellow-nosed Albatross {Diomedea chlororhynchos)<br />
and are referred to that species. Large numbers of<br />
these albatrosses currently breed on Amsterdam (Jouventin et<br />
al., 1984).<br />
Diomedea amsterdamensis<br />
The Martinez collection contained numerous remains of Diomedea<br />
amsterdamensis that were removed prior to THW's<br />
examination of the collection. These were reported on by Jouventin<br />
et al. (1989), who recorded bones representing 81 indi-<br />
TABLE 1.—List of species, number of bones, and minimum number of individuals<br />
(MNI) calculated as the sum of MNI in individual sites, as defined by the<br />
collectors, represented in the fossils from Amsterdam Island.<br />
Species<br />
Eudyptes chrysocome<br />
Diomedea sp.<br />
Diomedea amsterdamensis<br />
Diomedea chlororhynchos<br />
Procellaria cinerea<br />
Pterodroma macroptera<br />
Pterodroma mollis<br />
Pterodroma baraui<br />
Pterodroma baraui/arminjoniana<br />
Pterodroma arminjoniana<br />
Pterodroma sp. ?aterrima<br />
Pachyptila macgillivrayi<br />
Pachyptila cf. turtur<br />
Pachyptila cf. desolata<br />
Pachyptila sp. chicks<br />
Pelecanoides urinatrix<br />
Puffinus assimilis<br />
Puffinus carneipes<br />
Puffinus cf. griseus<br />
Pelagodroma marina<br />
Fregetta grallaria<br />
Oceanites sp.<br />
Hydrobatidae sp. indet.<br />
Catharacta skua hamiltoni<br />
Anas marecula<br />
Gallus gallus<br />
Total<br />
Number of bones<br />
281<br />
42<br />
80<br />
9<br />
980<br />
3711<br />
4766<br />
263<br />
11<br />
6<br />
2<br />
9235<br />
57<br />
1<br />
7<br />
106<br />
1882<br />
21<br />
2<br />
1780<br />
72<br />
17<br />
5<br />
179<br />
44<br />
3<br />
23562<br />
MNI<br />
21<br />
3<br />
9(8 ir<br />
2<br />
117<br />
282<br />
375<br />
31<br />
3<br />
4<br />
2<br />
731<br />
12<br />
1<br />
2<br />
29<br />
152<br />
1<br />
1<br />
206<br />
25<br />
4<br />
2<br />
19<br />
24(33)*<br />
1<br />
2059<br />
"Jouventin et al. (1989) recorded 81 individuals of D. amsterdamensis, and<br />
Olson and Jouventin (1996) reported a minimum of 33 individuals of Anas<br />
marecula, and there is probably overlap between those samples and what we<br />
examined in the study.<br />
viduals of D. amsterdamensis from several unspecified sites<br />
distributed between 80 m and 700 m a.s.l., including 52 individuals<br />
from a single site at 332 m, which is probably site 4.<br />
About 80 bones of a large species of albatross remained in the<br />
collection, however, and are referred to D. amsterdamensis because<br />
they do not match any other species. Diomedea amsterdamensis<br />
is restricted to breeding sites above 400 m (Jouventin<br />
etal., 1989).<br />
Pterodroma<br />
FIGURES 5-8<br />
A preliminary examination of the Pterodroma bones in the<br />
collection revealed the predominance of two different species<br />
that differed mainly in size. In order to characterize these better,<br />
a large sample was measured, revealing that both species<br />
have apparently normal distributions for all bones (Figures 7,<br />
8; Appendices 1,2).<br />
Species in the size range of the larger taxon include the<br />
Great-winged or Grey-faced Petrel {Pterodroma macroptera),<br />
Juan Fernandez Petrel {P. externa), Providence Petrel {P.<br />
solandri), Magenta Petrel {P. magentae), and White-necked