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30 SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY<br />

Cor. •<br />

Hum.<br />

Uln.<br />

Cpm. < i<br />

Fem.<br />

Tbt.<br />

Tmt.<br />

80<br />

I<br />

-100 120<br />

I<br />

140<br />

I<br />

160<br />

I<br />

180 200<br />

I<br />

• Psittacula krameri, standard<br />

* Mascarinus mascarinus, fossil<br />

D M. mascarinus, Recent, Paris and Vienna<br />

+ Necropsittacus rodericanus<br />

O Coracopsis nigra, min. and max.<br />

FIGURE 12.—Ratio-diagram of the dimensions of bones of fossil Mascarinus mascarinus from Reunion, compared<br />

with those taken from x-radiographs of the two mounted specimens (MNHN, Natural History Museum of<br />

Vienna) and with Coracopsis nigra. The standard is Psittacula krameri (Pierce Brodkorb 27712). For C. nigra<br />

the dimensions are the minimum and maximum of the specimens at AMNH (3571, 4399), MNHN (LAC 1883-<br />

507-1883-509), and USNM (224810, 292917, 432236). Coracoid measurement is of internal length; for other<br />

bones, measurement is of total length. When measurements are not known, successive points are united by<br />

dashed lines. (Cor.=coracoid, Cpm.=carpometacarpus, Fem.=femur, Hum.=humerus, Tbt.=tibiotarsus, Tmt.=<br />

tarsometatarsus, Uln.=ulna.)<br />

subspecies from the Comoros and Praslin are smaller (Forshaw,<br />

1973).<br />

According to the accounts of the early explorers, there were<br />

at least four species of parrots on Reunion. The description given<br />

by Dubois of "parrots a little bigger than pigeons, the feathering<br />

of the color of petit-gris, a black hood on the head, the<br />

beak very strong and the color of fire" is considered to apply to<br />

M. mascarinus (Barre and Barau, 1982:31, our translation).<br />

Petit-gris is the name given to the fur of the Eurasian Red<br />

Squirrel {Sciurus vulgaris Linnaeus) in its dark phase. Two<br />

other species were a grey parrot, which according to Feuilley<br />

was smaller than M. mascarinus, and a green parakeet with a<br />

black ring, the Reunion Ring-necked Parakeet, Psittacula<br />

eques (Boddaert), which probably was conspecific with the<br />

still-living Mauritian Echo Parakeet (Cheke, 1987). A fourth<br />

species, which was the same size as P. eques, was described by<br />

Dubois as a "green parrot with head, upper parts of wings, and<br />

tail color of fire" (Barre and Barau, 1982:31, our translation).<br />

FIGURE 13 (opposite).—Fossils of falcons, coots, pigeons, and parrots from the<br />

Mascarenes. Falco duboisi: a, left coracoid, Grotte de l'Autel, 330548, posterior<br />

view, xl.5; b, left tibiotarsus, paratype, Grotte des Premiers Francais, 1993-30,<br />

anterior view; c, right tibiotarsus, paratype, Grotte des Premiers Francais,<br />

1993-33, anterior view; d, left femur, paratype, Grotte des Premiers Francais,<br />

1993-29, posterior view; e, left tarsometatarsus, holotype, Grotte des Premiers<br />

Francais, 1993-28, anterior view. Falco punctatus: f left coracoid, Montagne du<br />

Pouce caves, Mauritius, MNHN, not numbered, posterior view, xl.5. Nesoenas<br />

duboisi: g, right humems, Grotte de l'Autel, 330546, anconal view; h, same, palmar<br />

view. Fulica newtonii: i, sternum, Grotte des Premiers Francais, 1993-39,<br />

ventral view;/ right tibiotarsus, Grotte des Premiers Francais, 199341, anterior<br />

view; k, right tarsometatarsus, Marais de l'Ermitage, 1811, anterior view; /, right<br />

coracoid, Marais de l'Ermitage, 1814, posterior view; m, left ulna, distal part,<br />

Marais de l'Ermitage, 1815, internal view. Streptopelia picturata: n, right<br />

humems, proximal part, Grotte "au sable," 330738, anconal view, xl.5. Mascarinus<br />

mascarinus: o, right humems, Grotte de l'Autel, 330539, anconal view;p,<br />

same, palmar view; q, left carpometacarpus, Grotte de l'Autel, 330544, internal<br />

view, xl.5; r, left coracoid, Grotte de l'Autel, 330545, posterior view, xl.5; 5,<br />

same, anterior view, xl.5; /, left ulna, Grotte de l'Autel, 330541, palmar view; u,<br />

same, internal view; v, left femur, Grotte de l'Autel, 330543, posterior view; w,<br />

same, anterior view; x, left tibiotarsus, Grotte de l'Autel, 330542, anterior view.<br />

Mascarenotus grucheti: y, right tarsometatarsus, holotype, Grotte des Premiers<br />

Francais, 199349, anterior view. (a,f n,q,r,s xl.5; others natural size.)

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