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280<br />

The left coracoid is represented by the shoulder end and<br />

shaft. The sternal end is covered by matrix and sternal bone,<br />

but it is well seen in the x-radiograph (Figure 5). The sternal<br />

end is wide and flat, with a long-pointed medial angle and with<br />

a rectangular lateral process. Such a structure of the medial angle<br />

is very similar to that in Lithornisplebius Houde, 1988. The<br />

acrocoracoid is sturdy, relatively short, and its dorsal top is<br />

three-edged and bluntly acute. The craniomedial side of the acrocoracoid<br />

bears an elongate depression that probably represents<br />

an articulation for the clavicle. The lateral side of the acrocoracoid<br />

possesses a wide, slightly concave depression of the<br />

acrocoracohumeral tendon. Ventral to this depression, a relatively<br />

small humeral articular facet is located, the facet being<br />

exposed laterally. An ellipsoidal scapular cotyla is exposed<br />

caudolaterally and is located on an enlarged base of a wide,<br />

flat, long procoracoid process. The sternal portion of the coracoidal<br />

shaft is strongly broadened. None of the elements of the<br />

shoulder girdle are compressed, and they all preserve the tme<br />

configurations of the bones.<br />

WING BONES.—The proximal end of the left humerus was<br />

strongly compressed in its plane during preservation. The humeral<br />

articular head is small, bean-shaped, and located in the<br />

ventral position of the proximal end. The humerus has a<br />

well-developed deltopectoral crest beginning very close to the<br />

humeral articular head in the most proximal position of the<br />

proximal end; it is similar to that in Lithornis plebius. The deltopectoral<br />

crest is flat but is rather deflected dorsally, contrary<br />

to Elzanowski (1995), who described it as projecting laterally.<br />

FIGURE 4.—Shoulder articulation in Ambiortus dementjevi on the opposite<br />

view of the main slab, PIN 3790-271+. ACR=acromion, APR=acrocoracoid<br />

process, ATB=acromial dorsal tubercle, CMR=caudal margin, DPT=dorsal<br />

pit, HAH=humeral artiular head, TRF=tricipital fossa, VTB=ventral tubercle.<br />

(Scale bar= 1 cm.)<br />

SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY<br />

FIGURE 5.—X-radiograph of the main slab with Ambiortus dementjevi. The<br />

sternal end of the coracoid with the medial angle and structure of the vertebrae<br />

can be clearly seen. CTP=caudal transverse process, LPC=lateral process of<br />

coracoid, MAN=medial angle of stemal end of coracoid, RB=rib, V10=10th<br />

vertebra.<br />

Along the dorsal margin, a shallow groove appears in the proximal<br />

half on the cranial side. The bicipital crest and pneumotricipital<br />

fossa are absent; the latter is expressed only as a tricipital<br />

depression (Figure 4). The ventral edge of the proximal end<br />

of the humerus is remarkably projected ventrally. Its distal<br />

edge is like a boss. The cranial surface of this boss possesses a<br />

slightly pronounced cranial tubercle with a pit in the center.<br />

Cranially from this tubercle is a noticeable ligamental fossa<br />

(not a groove). Lithornis plebius also has a similar tubercle<br />

possessing a pit and has a ligamental fossa instead of a furrow.<br />

Such a ligamental fossa is probably the homolog of the transverse<br />

ligamental furrow. On the caudal surface of the proximal<br />

end, a small dorsal pit is developed in the usual place of the<br />

dorsal tubercle. A small ventral tubercle is represented on the<br />

caudal surface of a projecting ventral edge. The capital groove<br />

is not developed. A slightly elevated caudal margin mns along<br />

the middle of the shaft and is directed toward the middle of the<br />

humeral head.<br />

The ulna is badly damaged. Only its distal end and a mold of<br />

a portion of the shaft are preserved on the main and associated

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