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276 concerning the amphicoelous structure of the vertebrae and the bones of the shoulder girdle. New preparation and observation of the specimen provide a complete and corrected anatomical description of this bird. It was a great surprise to discover that Ambiortus dementjevi is similar to Otogornis genghisi, which was described by Hou (1994) from the Ordos Basin at the Chabu Sumu locality, Otog Qi, Yikezhao-meng, Inner Mongolia, China. The specimen was collected in the thin, grey green mudstones of the Yijinhuoluo Formation of the Zidan Group in the Lycoptera-bearing deposits and represents the earliest Cretaceous or even a Late Jurassic avian fossil from China (Hou, 1994). Otogornis is based on associated elements of the forelimb and shoulder girdle (VP- 9607, holotype, Institute of Vertebrate Paleontology and Paleanthropology (IVPP), Beijing). The slab also displays some feather impressions. Otogornis was described as Aves incertae sedis and was compared with Archaeopteryx, Chaoyangia, and the enantiornithines Sinornis and Cathayornis (Hou, 1994). Earlier, the same specimen was assigned to the indeterminate Enantiornithes (Dong, 1993). ACKNOWLEDGMENTS.—I thank very much L. Hou and Z. Zhou for permitting investigation of the holotype of O. genghisi in the IVPP collection. L. Martin prompted me to use for the investigation the mold from specimen 3790-272, which led to the discovery of a contact with the counterslab specimen 3790-271- (specimens in the Paleontological Institute, Russian Academy of Sciences (PIN)). Comparison with the lifhornithids was made possible by the courtesy of R. Emry in the Department of Vertebrate Paleontology, National Museum of Natural History, Smithsonian Institution, Washington, D.C. For comments on the manuscript and English corrections I am thankful to S. Lucas, R. Zusi, S. Olson, and two anonymous reviewers. The stereophotographs were made by S. Morton in the Faculty of Sciences of Monash University, Clayton, Australia. The x-radiograph of Ambiortus dementjevi was made by L. Martin and J. Chom in the Museum of Natural History, University of Kansas. All drawings, including Otogornis, were made by the author. This study was supported by a travel grant and grant 96-04-50822 of the Russian Fund for Fundamental Research, as well as by funds from the IVPP, Academy of Sciences of China. Age of the Khurilt Beds The geological age of Ambiortus is problematical. It was found in Mesozoic rocks of the Gobian Altai in central Mongolia in the Khurilt-Ulan Bulak locality. This discovery caused some paleontologists to doubt the correct definition of the fossil and the age of the deposits. At present, no doubts exist about the advanced avian condition of this specimen. The geologic age of the Upper Cretaceous lacustrine shales and sandstones of the Khurilt locality, however, is discussed in contradictory terms by different experts in biostratigraphy, paleobotany, and SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY paleoentomology, with the dates ranging from the latest Jurassic to the Aptian. Numerous and various insects were collected in these beds (Zherikhin, 1978; Sinitsa, 1993). The insect fauna is very constant in a number of localities in central Mongolia (Khurilt, upper members of Kholbotoo, Bon Tsagan). This is known as the Bon Tsagan assemblage, the youngest among three Lower Cretaceous assemblages in central Mongolia (Ponomarenko, 1990). Dmitriev and Zherikhin (1988) supported an Aptian age of the deposits in these localities. The plant associations of the Khurilt, neighboring deposits of the Kholbotoo, and of the middle levels of the Bon Tsagan localities include, following Krassilov (1982), four phytostratigraphic units in the Upper Cretaceous of Mongolia. The third unit is the Baierella hastata (Bennettitales; and its cones, Karkenia mongolica)/'Araucaria mongolica zone, which includes localities of Shin Khooduk-Anda Khooduk level, and most of the paper shales of the Bon Tsagan, Kholbotoo Gol, Khurilt, Erdeni Ula, Shin Khooduk, and Modon Usoo localities. The sediments of the Khurilt and Kholbotoo localities were assigned by Krassilov (1982) to the Anda Khooduk Formation. The plant communities from these localities he correlated with the Aptian flora of the Russian Far East (Primorye). Thus, phytostratigraphic data suggest an Aptian age for the Khurilt beds (Krassilov, 1982). Based on geological data, Martinson (1973) and Shuvalov (1975, 1982, 1993) referred the Khurilt and Kholbotoo beds to the Anda Khooduk Formation, which they correlated with the Hauterivian-Barremian. Sinitsa (1993), based on the lithofacial data and ostracod assemblages, did not provide a definite age for the Khurilt beds and defined it as a task for future exploration. She specified that the Khurilt beds belong to the Khurilt Section of the Bon Tsagan Series. The Khurilt beds are underlaid by the Dund Argalant Series of the latest Jurassic, which includes the Anda Khooduk Formation (Tithonian), and are overlapped nearby by the sediments of the Kholbotoo Section (younger beds of the Bon Tsagan Series). In more western areas of Mongolia, the Bon Tsagan Series are overlapped by the Khoolsyn Gol Formation, which is correlated with the Aptian-Albian. Perhaps the Mongolian and Chinese Lower Cretaceous shale sediments were deposited simultaneously. The problem of age determination of the lacustrine Lower Cretaceous beds in Mongolia is the same as for the Jehol Group in China (Matsukawa and Obata, 1994). The Khurilt outcrop and the upper members of the neighboring Kholbotoo outcrop are very similar to the grey green, thin-bedded sandy and oil shales and siltstones of the Jiufotang Formation in Liaoning Province of China. The Jiufotang Formation is a member of the Jehol Group, which is subdivided into four lithostratigraphic units: Yixian, Jiufotang, Shahai, and Fuxin formations (Smith et al., 1995). The Jiufotang Formation is correlated with the Berriasian-Valanginian (Li and Liu, 1994), or Tithonian-Valanginian (Lin, 1994), or even with the Tithonian (Chen and Chang, 1994). On the basis
NUMBER 89 277 of fossil fishes, the Yixian and Jiufotang formations are correlated with the Late Jurassic and Neocomian, in agreement with the fish faunas of Japan, Kazakhstan, and western Europe (Fan, 1996). Published radiometric ages for the base of the Yixian Formation include 137±7 Ma using K/Ar and 142.5 Ma using Rb/Sr (Wang, 1983; Wang and Diao, 1984), which corresponds with a Berriassian age, using the time scale of Harland et al. (1989). For the Fuxin Formation, K/Ar ages range from 100 Ma to 137 Ma (Mao et al., 1990), corresponding with an Aptian-Valanginian age. New age dates were reported for the Yixian Formation, however, that are based on a laser 40 Ar/ 39 Ar study of single mineral grains (Smith et al., 1995). This study estimated the age of the lower Yixian Formation as 121.2±0.3 Ma and 121.3±2.3 and 121.4±0.7 Ma for the upper Yixian Formation. Smith et al. (1995) also tested the absolute ages using 40 Ar/ 39 Ar in very fine crystallites of glaucony from the white lacustrine Ershilipu sediments that occupy a stratigraphic position between the upper and lower parts of the Yixian Formation. The resulting ages of 122.1 ±0.2 Ma and 122.5±0.3 Ma agree with the strict chronostratigraphic constraints imposed by the 40 Ar/ 39 Ar ages of the upper and lower Yixian Formation. Thus, Smith et al. (1995) provided an integrated age range of 121.1-122.9 Ma for the Yixian Formation, which corresponds to the Barremian using the time scale of Harland et al. (1989), and these dates are much younger than the K/Ar and Rb/Sr dates of other authors. Thus, insects and plants suggest an Aptian age for the Khurilt deposits, but the geological data and ostracods indicate a Neocomian age. The probably contemporaneous Yixian and Jiufotang formations in northeastern China are assigned to the Neocomian on faunistic, plant, and radiometric data, although the laser 40 Ar/ 39 Ar study gives a Barremian age. I am inclined to accept a Neocomian age for the Khurilt deposits. Comprehensive Description of Ambiortus Ambiortus dementjevi is represented on three slabs. The main slab (PIN 3790-271+) bears the cervical and thoracic vertebrae, furcula, left scapula and coracoid, a portion of the sternum, some thoracic ribs, the proximal portion of the left humerus, distal portions of the radius and ulna, ulnare, the proximal portion of the left carpometacarpus, and phalanges of the major wing digit (Figures 1, 3-5). This slab also shows an isolated impression of a feather vane about 12 mm long and the probable impression of the soft body of the specimen, with small contour feathers that surround the body impression. The feathers and body impression are better represented on the counterslab (PIN 3790-271-). The counterslab bears small fragments of the vertebrae and a portion of the major metacarpal, and it has a good mold of the humems, coracoid, clavicle, and carpometacarpus. A small associated slab (PIN 3790-272) shows just the mold of three phalanges of the major left wing digit, the mold of the ulna and major metacarpal, and a frag- FlGURE 1.—Main slab with Ambiortus dementjevi Kurochkin, 1982, holotype PIN 3790-271+; Khurilt Ulan Bulak locality, central Mongolia, Neocomian. Stereopairs. (Scale bar= 1 cm.)
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NUMBER 89 277<br />
of fossil fishes, the Yixian and Jiufotang formations are correlated<br />
with the Late Jurassic and Neocomian, in agreement with<br />
the fish faunas of Japan, Kazakhstan, and western Europe (Fan,<br />
1996).<br />
Published radiometric ages for the base of the Yixian Formation<br />
include 137±7 Ma using K/Ar and 142.5 Ma using Rb/Sr<br />
(Wang, 1983; Wang and Diao, 1984), which corresponds with<br />
a Berriassian age, using the time scale of Harland et al. (1989).<br />
For the Fuxin Formation, K/Ar ages range from 100 Ma to 137<br />
Ma (Mao et al., 1990), corresponding with an Aptian-Valanginian<br />
age. New age dates were reported for the Yixian Formation,<br />
however, that are based on a laser 40 Ar/ 39 Ar study of single<br />
mineral grains (Smith et al., 1995). This study estimated the<br />
age of the lower Yixian Formation as 121.2±0.3 Ma and<br />
121.3±2.3 and 121.4±0.7 Ma for the upper Yixian Formation.<br />
Smith et al. (1995) also tested the absolute ages using 40 Ar/<br />
39 Ar in very fine crystallites of glaucony from the white lacustrine<br />
Ershilipu sediments that occupy a stratigraphic position<br />
between the upper and lower parts of the Yixian Formation.<br />
The resulting ages of 122.1 ±0.2 Ma and 122.5±0.3 Ma agree<br />
with the strict chronostratigraphic constraints imposed by the<br />
40 Ar/ 39 Ar ages of the upper and lower Yixian Formation. Thus,<br />
Smith et al. (1995) provided an integrated age range of<br />
121.1-122.9 Ma for the Yixian Formation, which corresponds<br />
to the Barremian using the time scale of Harland et al. (1989),<br />
and these dates are much younger than the K/Ar and Rb/Sr<br />
dates of other authors.<br />
Thus, insects and plants suggest an Aptian age for the Khurilt<br />
deposits, but the geological data and ostracods indicate a<br />
Neocomian age. The probably contemporaneous Yixian and Jiufotang<br />
formations in northeastern China are assigned to the<br />
Neocomian on faunistic, plant, and radiometric data, although<br />
the laser 40 Ar/ 39 Ar study gives a Barremian age. I am inclined<br />
to accept a Neocomian age for the Khurilt deposits.<br />
Comprehensive Description of Ambiortus<br />
Ambiortus dementjevi is represented on three slabs. The<br />
main slab (PIN 3790-271+) bears the cervical and thoracic vertebrae,<br />
furcula, left scapula and coracoid, a portion of the sternum,<br />
some thoracic ribs, the proximal portion of the left humerus,<br />
distal portions of the radius and ulna, ulnare, the<br />
proximal portion of the left carpometacarpus, and phalanges of<br />
the major wing digit (Figures 1, 3-5). This slab also shows an<br />
isolated impression of a feather vane about 12 mm long and the<br />
probable impression of the soft body of the specimen, with<br />
small contour feathers that surround the body impression. The<br />
feathers and body impression are better represented on the<br />
counterslab (PIN 3790-271-). The counterslab bears small fragments<br />
of the vertebrae and a portion of the major metacarpal,<br />
and it has a good mold of the humems, coracoid, clavicle, and<br />
carpometacarpus. A small associated slab (PIN 3790-272)<br />
shows just the mold of three phalanges of the major left wing<br />
digit, the mold of the ulna and major metacarpal, and a frag-<br />
FlGURE 1.—Main slab with Ambiortus dementjevi Kurochkin, 1982, holotype PIN 3790-271+; Khurilt Ulan<br />
Bulak locality, central Mongolia, Neocomian. Stereopairs. (Scale bar= 1 cm.)