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18 SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY
odori as well as in A. gibberifrons. The humerus of A. theodori
also looks very similar to that of A. gibberifrons. The humeral
head forms a well-developed rim above the tricipital fossa, and
the pectoral attachment is well developed and elongated. On the
distal part, the brachialis anticus impression and the olecranal
fossa are deep. The ulna of A. theodori (Figure Iq) is short and
stout, exactly as in A. gibberifrons, and its size is the same as in
a male of that species (USNM 610562). The humero-ulnar depression
is pronounced proximally. Distally, the external
condyle is extended by a narrow and well-defined lip, which rises
proximad along the shaft, as in A. gibberifrons. The tibiotarsus
and tarsometatarsus, compared with those of A. gibberifrons,
are longer, more robust, and their proximal and distal parts are
proportionally wider (Table 7).
The proportions of A. theodori are very close to those of A.
gibberifrons (Figure 6), but the coracoid, tibiotarsus, and tar-
Cor.
Hum.
Uln.
Cpm.
Fern.
Tbt.
Tmt.
• Anas hottentota, standard
• Anas marecula
)k Anas bernieri
A Anas aucklandica
+ Anas gibberifrons cj
O Anas theodori
FlGURE 7 (opposite).—Fossils of ibis and waterfowl from Reunion Island.
Threskiornis solitarius: a, left tibiotarsus, Marais de l'Ermitage, 1867, anterior
view; b, same, posterior view; c, right tarsometatarsus, Marais de
1'Ermitage, 1801, anterior view; d, same, posterior view; e, right tarsometatarsus,
distal part, Grotte des Premiers Francais, 1993-37, anterior view. Alopochen
(Mascarenachen) kervazoi: f, cranium, Grotte de l'Autel, 330525,
forsal view; g, rostrum, holotype, Grotte des Premiers Francais, 1993-19,
dorsal view; h, left coracoid, Grotte de l'Autel, 330519, posterior view; i,
right humerus, Grotte de l'Autel, 330517, anconal view;y, right tarsometatarsus,
Grotte de l'Autel, 330521, anterior view; k, right carpometacarpus,
Grotte de l'Autel, 330522, internal view; /, right carpometacarpus, paratype,
Grotte des Premiers Francais, 1993-20, internal view; m, left femur, Grotte de
l'Autel, 330526, posterior view; n, sternum, anterior part, Grotte de l'Autel,
330523, right lateral view. Aythya sp.: o, right carpometacarpus, Marais de
l'Ermitage, 1924, internal view; p, left carpometacarpus, proximal part,
Marais de l'Ermitage, 1925, internal view. Anas theodori: q, left ulna, Marais
de l'Ermitage, 1810, internal view; r, left tibiotarsus, Marais de l'Ermitage,
1894, anterior view. All figures are natural size.
-150 -100 -50 50 100 150
!
FIGURE 6.—Ratio-diagram of the mean dimensions of Anas theodori compared with A. gibberifrons (male,
USNM 610562) and to the mean dimensions of two Anas bernieri (Steve Goodman, pers. comm., 1995). The
standard is Anas hottentota (male, USNM 430832). For comparison, the dimensions of two flightless forms are
indicated, Anas aucklandica (USNM 612796), and the extinct Anas marecula, after Martinez (1987). For A.
bernieri the internal length of the coracoid has been estimated from its total length. Coracoid measurement is of
internal length; for other bones, measurement is of total length. When measurements are not known, successive
points are united by dashed lines. (Cor.=coracoid, Cpm.=carpometacarpus, Fem.=femur, Hum.=humerus
Tbt.=tibiotarsus, Tmt.=tarsometatarsus, Uln.=ulna.)
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