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NUMBER 89 17<br />

cal in shape and dimensions to that from Mauritius and therefore<br />

must belong to the same species.<br />

Compared with living forms of Anas from Madagascar, A.<br />

theodori is larger than A. bernieri (Hartlaub) (Figure 5, Table 7)<br />

and smaller than A. melleri P.L. Sclater (Howard, 1964). We<br />

compared it in the USNM with various species of Anas. The one<br />

it resembles most is Anas gibberifrons Muller, which lives in the<br />

east and northeast of the Indian Ocean, from the Andaman Islands<br />

to Indonesia and Australia, and on islands in the western<br />

Pacific Ocean (Mayr and Cottrell, 1979). The common characteristics<br />

and the differences between A. theodori and A. gibberi­<br />

Measurement<br />

Sternum<br />

width (a)<br />

keel depth (b)<br />

Coracoid<br />

internal length<br />

proximal width<br />

proximal depth<br />

stemal-facet length<br />

stemal-facet depth<br />

minimum shaft width<br />

depth at same level<br />

Humerus<br />

total length<br />

proximal width<br />

proximal depth<br />

distal width<br />

distal depth<br />

midshaft width<br />

midshaft depth<br />

Ulna<br />

total length<br />

proximal width<br />

proximal depth<br />

distal width<br />

depth external condyle<br />

midshaft width<br />

midshaft depth<br />

Tibiotarsus<br />

total length (c)<br />

proximal width (c)<br />

proximal depth (c)<br />

distal width<br />

distal depth<br />

minimum shaft width<br />

depth at same level<br />

Tarsometatarsus<br />

total length<br />

proximal width<br />

proximal depth<br />

distal width<br />

distal depth<br />

midshaft width<br />

midshaft depth<br />

'Specimen very juvenile.<br />

frons are as follows. If the anterior part of the sternum illustrated<br />

by Newton and Gadow (1893, pl. 34: figs. 11, 12) is perfectly<br />

preserved, the carina projects further anteriorly and the ventral<br />

manubrial spine (spina externa of Newton and Gadow) is narrow<br />

and elongated in A. gibberifrons and is more elongated than in A.<br />

theodori. The coracoid of A. theodori looks very similar to that<br />

of A. gibberifrons. On the internal side of the posterior face,<br />

above the sternal facet, there is a small, proximally directed<br />

spike, followed by a small depression, which also is present in A.<br />

gibberifrons. On the anterior face, near the internal side of the<br />

acrocoracoid, an almost circular muscular scar occurs in A. the-<br />

TABLE 7.—Dimensions (mm) of the main bones of the extinct Anas theodori, from Mauritius and Reunion, compared<br />

with modem Anas gibberifrons (USNM) and Anas bernieri (two specimens; Steve Goodman, pers. comm.,<br />

1995). For Anas theodori from Mauritius, the dimensions are from material at MNHN (prefix MAD), from Newton<br />

and Gadow (N. & G., 1893), or measured from their figures; Reunion dimensions are from material at<br />

MHNR (prefix FE-O). (a=width of front part of sternum, measured between the two lateral intermuscular lines;<br />

b=stemum-keel depth, from dorsal surface of sternum to ventral tip of keel; c=measured without cnemial crests;<br />

d=measured with cnemial crests; «=number of specimens.)<br />

Reunion<br />

FE-O-1810<br />

-63<br />

7.5<br />

7.0<br />

6.0<br />

7.1<br />

4.4<br />

3.9<br />

FE-O-1894<br />

71.0<br />

8.5<br />

9.7<br />

8.0<br />

-<br />

3.8<br />

3.0<br />

Mauritius<br />

MAD-4057 (cast)<br />

27.7<br />

21.7<br />

MAD-4993<br />

41.1<br />

-9<br />

8.3<br />

-16<br />

3.6<br />

4.3<br />

3.5<br />

MAD-4988<br />

73.0<br />

-16<br />

-<br />

11.2<br />

6.6<br />

5.2<br />

4.5<br />

MAD-7161<br />

71.0<br />

8.6<br />

9.2<br />

8.0<br />

9.0<br />

3.7<br />

3.2<br />

MAD-4992*<br />

-42<br />

-<br />

-<br />

-<br />

4.2<br />

3.6<br />

Anas theodori<br />

Mauritius<br />

N. &G., 1893<br />

42.5<br />

-8<br />

-<br />

17.2<br />

-<br />

4.1<br />

-<br />

MAD-4989<br />

69.6<br />

15.5<br />

7.9<br />

11.3<br />

6.5<br />

5.2<br />

4.2<br />

N. &G., 1893<br />

42.0<br />

9.2<br />

8.3<br />

9.2<br />

-<br />

4.3<br />

-<br />

Mauritius<br />

MAD-4991<br />

73.1<br />

16.1<br />

8.2<br />

11.6<br />

6.5<br />

5.6<br />

4.6<br />

Anas bernieri<br />

-30.9,-31.2<br />

-<br />

-<br />

-<br />

-<br />

-<br />

64.5, 66.5<br />

13.3, 13.5<br />

-<br />

10.0, 10.3<br />

- --<br />

58.0, 60.6<br />

7.4, 7.6<br />

-<br />

6.0,6.1<br />

-<br />

-<br />

65.1, 67.6(d)<br />

6.7, 6.8<br />

-<br />

6.7, 6.8<br />

-<br />

-<br />

-<br />

38.7, 39.3<br />

7.0, 7.3<br />

-<br />

7.3, 7.5<br />

-<br />

-<br />

mean (n=4)<br />

24.25<br />

20.78<br />

37.55<br />

8.03<br />

7.20<br />

15.33<br />

2.98<br />

3.80<br />

2.98<br />

69.93<br />

15.18<br />

7.68<br />

10.75<br />

6.50<br />

5.33<br />

4.50<br />

61.55<br />

7.45<br />

6.88<br />

5.98<br />

6.50<br />

3.95<br />

4.00<br />

64.38<br />

7.50<br />

8.53<br />

6.78<br />

7.58<br />

3.08<br />

2.68<br />

36.15<br />

7.48<br />

6.83<br />

7.70<br />

5.48<br />

3.35<br />

2.93<br />

Anas gibberifrons<br />

range<br />

23.2-25.8<br />

19.4-21.8<br />

35.1-38.9<br />

7.4-8.4<br />

6.8-7.6<br />

14.5-16.3<br />

2.7-3.2<br />

3.7^t.O<br />

2.6-3.2<br />

-68-71.4<br />

14.5-15.5<br />

7.4-8.0<br />

9.9-11.2<br />

6.2-6.7<br />

5.1-5.6<br />

4.2^1.8<br />

59.5-63.2<br />

7.1-7.6<br />

6.2-7.5<br />

5.7-6.1<br />

6.2-6.8<br />

3.6-^.2<br />

3.6-4.4<br />

59.5-67.4<br />

-7-7.9<br />

-8-9.3<br />

6.6-7.1<br />

7.0-8.0<br />

3.0-3.1<br />

2.5-2.8<br />

33.3-38.9<br />

7.1-8.1<br />

6.3-7.2<br />

7.1-8.3<br />

5.1-5.9<br />

3.0-3.9<br />

2.7-3.1

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