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228<br />

SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY<br />

FIGURE 8.—The left femur of Chaunoides antiquus, n. gen. n. sp. (paratype MNRJ 4621 -V), coated with ammonium<br />

chloride (right) compared with Chauna chavaria (USNM 347738) (left): A, anterior view; B, posterior<br />

view; c, distal view. (Scale bar= 1 cm.)<br />

shaft. In the proximal end of the tarsometatarsus (Figure 10),<br />

the intercotylar prominence is higher and sharper than in other<br />

Anhimidae; the morphology of the hypotarsus is very similar to<br />

that in Chauna, especially in C. chavaria, but the crista medialis<br />

is shorter and higher. Also, there is no pneumatic foramen in<br />

the proximal end of the tarsometatarsus, whereas in extant anhimids<br />

this region is very pneumatized.<br />

Discussion<br />

The procoracoid foramen in Chaunoides is large and perfectly<br />

formed, as it is in the available specimens of Anhima and<br />

some specimens of Chauna; however, in certain specimens,<br />

such as Chauna chavaria (USNM 347738) and C. torquata<br />

(HA 389), this foramen is not formed (Figure 5), a condition<br />

possibly due to immaturity or to intraspecific variation. Olson<br />

(1987) commented on this same variation in several genera of<br />

Accipitridae as possible intraspecific or intrageneric variation.<br />

The pneumatic foramen in the sterno-coracoidal fossa of<br />

Chaunoides and Chauna is a very large opening with rounded<br />

borders, being the extreme form of a condition that also is<br />

found in Anhima, Anseranas, Opisthocomus, gruids, some galliforms,<br />

and cathartids. In the three species of living anhimids,

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