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194 SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY TABLE 3.—Occurrence of mammalian taxa of presumed South American origin, and birds with Neotropical and western affinities, in faunas of late Pliocene to early Pleistocene (late Blancan to early Irvingtonian; 3.7-1.0 Ma) age in western North America and the Florida peninsula. References used to compile this table include Akersten (1972), Carranza-Castaneda and Miller (1988), Conrad (1980), Dalquest (1975), Downs and White (1968), Emslie (1988, 1992, 1995, 1998), Frazier (1981), Galusha et al. (1984), Gillette and Ray (1981), Hager (1974), Hirschfeld and Webb (1968), Hulbert (1992, 1997), Hulbert and Morgan (1993), Jefferson (1989), Johnson et al. (1975), Johnston and Savage (1955), Lindsay (1978, 1984a, 1984b), Lindsay and Tessman (1974), Lundelius et al. (1987), Miller and Carranza-Castaneda (1984), Montellano-Ballesteros and Carranza-Castaneda (1986), Morgan (1991), Morgan and Hulbert (1995), Opdyke et al. (1977), Robertson (1976), Schultz (1977, 1990), Schultz (1937), Seymour (1993), Skinner and Hibbard (1972), Tomida (1987), and Webb and Wilkins (1984). Inferred general habitat requirements for each taxon is indicated by superscript numbers as follows: 'thorn-scrub and savannah, Rowland tropical forest and/or hammock, and 3 aquatic or semiaquatic. Habitat assignment is based on that of living counterparts and/or paleoecological and paleobiological information provided in American Ornithologists' Union (1998), Brown and Amadon (1968), Campbell and Tonni (1981), Delacour and Amadon (1973), Downing and White (1995), Kurten and Anderson (1980), McDonald (1995), and Steadman (1980). (AZ=Arizona, CA=Califomia, CO=Colorado, ID=Idaho, MX=Mexico, TX=Texas.) Taxon Mammals Dasypus bellus' Holmesinafloridanus' Glyptotherium arizonae 3 Glyptotherium texanum 3 Pachyarmatherium leiseyi '• 2 Glossotherium chapadmalense' Glossotherium garbanii' Glossotherium sp. 1 Paramylodon harlani' Megalonyx leptostomus 2 Megalonyx wheatleyi 2 Megalonyx sp. 2 Eremotherium sp. 2 Nothrotheriops texanus Nothrotheriops sp. 1 Myrmecophaga tridactyla' Erethizon bathygnathum 2 Erethizon kleini 2 Erethizon dorsatum Erethizon poyeri 2 Neochoerus dichroplax 2,3 Neochoerus cordobai 2 ' 3 Neochoerus sp. 2 ' 3 Hydrochaeris holmesi 2 ' 3 Birds Phalacrocorax idahensis* Teratornis incredibilis^ 3 Teratornis merriami 1 - 3 Gymnogyps kofordi' Gymnogyps sp.' Neophrontops slaughteri' Amplibuteo concordatus '• 2 Aquila bivia' Spizaetus sp. 2 Cracidae, indet. 1 Meleagris leopoldi/anza 1 ' 2 Titanis walleri y Glaucidium explorator^ 2 Glaucidium sp. 1,2 = := o ac x Florida Western North America +* •- 3 ~ 75 (* N < Knolls, N < P> (IN O e —. < ^ £ ? •s OH a H= X J X X X X X X X x x x x x x x x X X X X X X X X X X X X X X X X X X X X X X a Wolf Ranch, California Wash, Cal Tech, Benson, Mendevil Ranch, McRae Wash, Curtis Ranch b Upper Arroyo Seco, lower Vallecito Creek < U X 1, T :d Corra * speth, T CO T3 X X X X X X X X S ta Clara X 2 ilancan' ,CO Q igerman

NUMBER 89 195 FIGURE 6.—Distribution of late Blancan to early Irvingtonian (ca. 2.5-1.0 Ma) local faunas in North America. Size of dot or circle indicates number of species of mammals of South American origin (Xenarthra, Caviomorpha) in the fauna. Local faunas include those listed in Table 3 plus Wellsch Valley, Saskatchewan; Delmont, South Dakota; Big Springs, Nebraska; Kentuck, Kansas; and Anita, Arizona. Arrows signify likely corridors of dispersal; the Gulf of Mexico terrestrial corridor would have been widened onto the continental shelf during glacial periods. these taxa suggests that the Gulf Coast corridor was composed of a mosaic of communities, including dry thorn-scrub, hammocks, and aquatic zones (lakes and wetlands). Such a broad zone having patches of dry to moist habitats is unlike any such region today and probably developed in response to unusual climatic conditions during glacial intervals in the Plio-Pleistocene. Conclusion The record of birds in North America during the Great American Biotic Interchange indicates patterns for timing and dispersals that are similar to those known for other vertebrates and for plants. As with the mammals, the birds are largely presumed to represent a xeric, thorn-scrub and savannah envi-

194 SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY<br />

TABLE 3.—Occurrence of mammalian taxa of presumed South American origin, and birds with Neotropical and western affinities,<br />

in faunas of late Pliocene to early Pleistocene (late Blancan to early Irvingtonian; 3.7-1.0 Ma) age in western North America and<br />

the Florida peninsula. References used to compile this table include Akersten (1972), Carranza-Castaneda and Miller (1988), Conrad<br />

(1980), Dalquest (1975), Downs and White (1968), Emslie (1988, 1992, 1995, 1998), Frazier (1981), Galusha et al. (1984),<br />

Gillette and Ray (1981), Hager (1974), Hirschfeld and Webb (1968), Hulbert (1992, 1997), Hulbert and Morgan (1993), Jefferson<br />

(1989), Johnson et al. (1975), Johnston and Savage (1955), Lindsay (1978, 1984a, 1984b), Lindsay and Tessman (1974), Lundelius<br />

et al. (1987), Miller and Carranza-Castaneda (1984), Montellano-Ballesteros and Carranza-Castaneda (1986), Morgan (1991), Morgan<br />

and Hulbert (1995), Opdyke et al. (1977), Robertson (1976), Schultz (1977, 1990), Schultz (1937), Seymour (1993), Skinner<br />

and Hibbard (1972), Tomida (1987), and Webb and Wilkins (1984). Inferred general habitat requirements for each taxon is indicated<br />

by superscript numbers as follows: 'thorn-scrub and savannah, Rowland tropical forest and/or hammock, and 3 aquatic or<br />

semiaquatic. Habitat assignment is based on that of living counterparts and/or paleoecological and paleobiological information<br />

provided in American Ornithologists' Union (1998), Brown and Amadon (1968), Campbell and Tonni (1981), Delacour and Amadon<br />

(1973), Downing and White (1995), Kurten and Anderson (1980), McDonald (1995), and Steadman (1980). (AZ=Arizona,<br />

CA=Califomia, CO=Colorado, ID=Idaho, MX=Mexico, TX=Texas.)<br />

Taxon<br />

Mammals<br />

Dasypus bellus'<br />

Holmesinafloridanus'<br />

Glyptotherium arizonae 3<br />

Glyptotherium texanum 3<br />

Pachyarmatherium leiseyi '• 2<br />

Glossotherium chapadmalense'<br />

Glossotherium garbanii'<br />

Glossotherium sp. 1<br />

Paramylodon harlani'<br />

Megalonyx leptostomus 2<br />

Megalonyx wheatleyi 2<br />

Megalonyx sp. 2<br />

Eremotherium sp. 2<br />

Nothrotheriops texanus<br />

Nothrotheriops sp. 1<br />

Myrmecophaga tridactyla'<br />

Erethizon bathygnathum 2<br />

Erethizon kleini 2<br />

Erethizon dorsatum<br />

Erethizon poyeri 2<br />

Neochoerus dichroplax 2,3<br />

Neochoerus cordobai 2 ' 3<br />

Neochoerus sp. 2 ' 3<br />

Hydrochaeris holmesi 2 ' 3<br />

Birds<br />

Phalacrocorax idahensis*<br />

Teratornis incredibilis^ 3<br />

Teratornis merriami 1 - 3<br />

Gymnogyps kofordi'<br />

Gymnogyps sp.'<br />

Neophrontops slaughteri'<br />

Amplibuteo concordatus '• 2<br />

Aquila bivia'<br />

Spizaetus sp. 2<br />

Cracidae, indet. 1<br />

Meleagris leopoldi/anza 1 ' 2<br />

Titanis walleri y<br />

Glaucidium explorator^ 2<br />

Glaucidium sp. 1,2<br />

= := o<br />

ac x<br />

Florida Western North America<br />

+* •- 3 ~<br />

75 (*<br />

N<br />

<<br />

Knolls,<br />

N<br />

<<br />

P> (IN O e —. < ^ £ ?<br />

•s OH a H=<br />

X J<br />

X X X X X X X<br />

x x x x x x x x<br />

X X X X X<br />

X X X X<br />

X X X<br />

X X<br />

X X X<br />

X<br />

X<br />

X<br />

X X<br />

a Wolf Ranch, California Wash, Cal Tech, Benson, Mendevil Ranch, McRae Wash, Curtis Ranch<br />

b Upper Arroyo Seco, lower Vallecito Creek<br />

<<br />

U<br />

X<br />

1, T<br />

:d Corra<br />

*<br />

speth, T<br />

CO T3<br />

X X X<br />

X X<br />

X X<br />

X<br />

S<br />

ta Clara<br />

X<br />

2<br />

ilancan'<br />

,CO<br />

Q<br />

igerman

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