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172 SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY<br />

FIGURE 2.—Map showing location of locality SBCM 05.006.399 for the new<br />

partial humerus referred to Aiolornis incredibilis. Dashed line indicates unsurveyed<br />

boundary of Temecula Land Grant.<br />

transverse section through nutrient foramen on margo ventralis<br />

at midshaft a flattened oval not quite twice as wide as deep<br />

(more egg-shaped, with margo ventralis slightly narrower than<br />

margo dorsalis in Argentavis; nearly round in Teratornis) (Figure<br />

3); (4) corpus humeri less curved proximally in both horizontal<br />

and vertical planes; and (5) attachment of M. pectoralis<br />

with distal portion a broad, flattened area (area damaged but<br />

appears similar in Argentavis; narrow, steeply sloping on distal<br />

end of crista deltopectoralis in Teratornis). These characters<br />

hold in comparison with all humeri of Teratornis from Rancho<br />

La Brea, California (n=>50).<br />

Although the position of the M. latissimus dorsi is so markedly<br />

different from that seen in known teratorns, even to the<br />

oooO<br />

A B C D<br />

FIGURE 3.—Comparison of the size and shape of the cross section of the humeral<br />

shaft in the plane of the nutrient foramen in a vulturid (A) and teratorns (B-D): A,<br />

California Condor, Gymnogyps californianus; B, Merriam's Teratorn, Teratornis<br />

merriami; C, Aiolornis incredibilis; D, Argentavis magnificens. Scale bar=2 cm.<br />

point of leading one to question whether or not the humerus<br />

might even be from a new, unknown family, in the absence of<br />

additional characters the presence of the four teratorn autapomorphies<br />

listed above are sufficient grounds for placing the<br />

specimen with the teratorns.<br />

Measurements of this specimen are limited to shaft width<br />

and depth in the plane of the midshaft nutrient foramen on the<br />

facies ventralis: 37.8 mm and 22.0±1 mm, respectively. Comparable<br />

measurements for the single specimen of Argentavis<br />

are 52.2 mm and 32.8 mm, respectively, and measurements for<br />

Teratornis merriami {n=\6) are 23.2-28.9 mm (x=24.9 mm)<br />

and 18.3-23.0 mm (JC = 19.9 mm), respectively.<br />

Right radius: Distal end, Anza-Borrego Desert State Park<br />

Paleontological Collection, ABDSP(LACM) 1318/V3803, from<br />

Irvingtonian (middle Pleistocene, or 1.5-1.0 Ma (Savage and<br />

Curtis, 1970:223)) deposits at the 3600-ft level of the Vallecito<br />

Creek area, Anza-Borrego Desert State Park, referred to Teratornis<br />

incredibilis by Howard (1963). Emended description, in<br />

comparison with T merriami, as follows: distal contour straighter,<br />

less rounded at corners; ligamental prominence extending<br />

more proximad, significantly elevated above and more markedly<br />

set off from shaft and more elevated than central ligamental attachment<br />

in anterior view (elevational differences slightly magnified<br />

by loss of surface bone on central ligamental attachment;<br />

both areas of attachment at about same level in Teratornis)<br />

(bone slightly crushed along proximal edge of prominence, but<br />

this cannot account for sharp drop to shaft); shaft immediately<br />

above distal end flat in anterior view; internal edge of shaft proximal<br />

to internal end of scapholunar facet broad, channeled, or<br />

slightly convex transversely, which gives way to knife-like ridge<br />

proximad, with interno-anterior edge formed by long, prominent<br />

attachment of Lig. radioulnare interosseum +Lig. ulno-radiocarpale<br />

(edge very narrow in Teratornis, sloping steeply mediad<br />

with attachment of Lig. radioulnare interosseum+Lig. ulno-radiocarpale<br />

a broad, flat area halfway between edge and midline<br />

of bone that is slightly elevated proximally); depression occurs<br />

between attachment of Lig. radioulnare interosseum + Lig. ulnoradiocarpale<br />

and midline of bone (accentuated, but not caused by<br />

crushing) (absent in Teratornis); central ligamental prominence<br />

more elevated and extending closer to distal end before dropping<br />

off than in Teratornis; external edge of shaft proximal to ligamental<br />

prominence with linear convexity; and shaft significantly<br />

more curved in anterior view and slightly more curved in external<br />

view (some of the curvature, but not much, may be a result of<br />

distortion in preservation).<br />

Premaxillary: Anza-Borrego Desert State Park Paleontological<br />

Collection, ABDSP(LACM) 6747/V26697, from Blancan<br />

(late Pliocene, or 3.5-3.2 Ma (Savage and Curtis,<br />

1970:223; Evernden et al., 1964:164)) deposits at the<br />

7000-8000-ft level of the Fish Creek beds in the Anza-Borrego<br />

Desert State Park, referred to Teratornis incredibilis by<br />

Howard (1972).<br />

Emended description as follows: markedly compressed, deep<br />

beak; palatal surface fairly flat (more concave than in Terator-

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