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NUMBER 89 155<br />
the period studied. This is probably a safe assumption for most<br />
of the seabirds in this study. It should be noted that none of the<br />
31 faunas contains seabird species with allopatric ranges (although<br />
this might have been the case if Puffinus yelkouan had<br />
been treated as a distinct species). This absence of allopatric<br />
species is in marked contrast to late Pleistocene nonmarine avifaunas,<br />
where species that are today widely allopatric (e.g.,<br />
Nyctea scandiaca and Alectoris spp.) frequently occur together.<br />
This similarity with modern seabird faunas also is an argument<br />
that the ecology of the species concerned has remained<br />
fairly constant over the period studied.<br />
Most of the reasonably well-dated faunas described above<br />
are either from the western Mediterranean or from Norway.<br />
The western Mediterranean faunas mostly suggest cool, biologically<br />
productive seas, and the western Mediterranean together<br />
with the waters around the Macaronesian islands may<br />
have been an important refugium for boreal seabirds during<br />
the coldest parts of the glaciation. Unfortunately, there is a<br />
dearth of sites from the mildest parts of the glaciation and<br />
Alcover, J.A., F. Florit, C. Mourer-Chauvire, and P.D.M. Weesie<br />
1992. The Avifaunas of the Isolated Mediterranean Islands during the<br />
Middle and Late Pleistocene. In K.E. Campbell, editor, Papers In<br />
Avian Paleontology Honoring Pierce Brodkorb. Science Series, Natural<br />
History Museum of <strong>Lo</strong>s Angeles County, 36:273-284.<br />
Alcover, J.A., S. Moya-Sola, and J. Pons-Moya<br />
1981. Les quimeres del passat. 260 pages. Ciutat de Mallorca: Editorial<br />
Moll. [Instirucio Catalana d'Histdria Natural filial de l'lnstitut<br />
d'Estudis Catalans, Memoria 11.]<br />
Ascenzi, A., and A.G. Segre<br />
1971a. A New Neandertal Child Mandible from an Upper Pleistocene Site<br />
in Southern Italy. Nature, <strong>Lo</strong>ndon, 233(5317):280-283.<br />
1971b. II giacimento con mandibola neandertaliana di Archi (Reggio Calabria).<br />
Accademia Nazionale dei Lincei, Rendiconti della Classe di<br />
Scienze Fisiche, Matematiche e Naturali, series 8, 50(6):763-771.<br />
Bell, A.<br />
1922. Pleistocene and Later Birds of Great Britain and Ireland. Naturalist,<br />
Hull, 1922(Aug-Sept):251-253.<br />
Bengtson, S.-A.<br />
1984. Breeding Ecology and Extinction of the Great Auk (Pinguinus impennis):<br />
Anecdotal Evidence and Conjectures. Auk, 101:1-12.<br />
Bietti, A.<br />
1987. Some Remarks on the New Radiocarbon Dates from the Arene Candide<br />
Cave (Savona, Italy). Human Evolution, 2(2): 185-190.<br />
Boessneck, J., and A. von den Driesch<br />
1980. Tierknochenfunde aus der sudspanischen hohlen. Studien uber<br />
Fruhe Tierknochenfunde von der Iberischen Halbinsel, 7:160-185.<br />
Bowen, D.Q.<br />
1970. The Palaeoenvironment of the 'Red Lady' of Paviland. Antiquity,<br />
44:134-136.<br />
Cassoli, P.F.<br />
1980. L'Avifauna de pleistocene superiore delle Arene Candide (Liguria).<br />
Memorie dell'Istituto Italiano di Paleontologia Umana, new series,<br />
3:155-234.<br />
Literature Cited<br />
from the eastern Mediterranean basin, although an isolated<br />
record shows that Morus bassanus occurred as far east as<br />
Crete during some part of the last glaciation (Suriano,<br />
1980).<br />
The Norwegian faunas, of course, date only from the milder<br />
parts of the glaciation and are perhaps most interesting as illustrations<br />
of violent climatic and environmental shifts during the<br />
late Pleistocene. This is particularly striking when comparing<br />
the Blomvag fauna (Figure 9) from the mild Boiling interstadial<br />
with the 2000 years younger Grotta Romanelli fauna (Figure<br />
11) from the cold Dryas 3 stadial. The occurrence of Pinguinus<br />
impennis in both these faunas is particularly noteworthy and<br />
suggests that even this nonvolant species was capable of<br />
changing its distribution quite rapidly in response to changing<br />
conditions. This contradicts Bengtson's (1984) theory that the<br />
extinction of Pinguinus impennis was at least partly due to an<br />
inability to adapt to environmental changes and supports the<br />
traditional view that the species' extinction was directly caused<br />
by human action.<br />
Cassoli, P.F., and A.G. Segre<br />
1985. L'Alca impenne Alca impennis del Pleistocene d'ltalia. In M. Fasola,<br />
editor, Atti 111 Convegno Italiano di Ornitologia, pages<br />
251-254.<br />
Cassoli, P.F., A.G. Segre, and E. Segre<br />
1979. Evolution morphologique et ecologique de la cote de Castro (Pouilles)<br />
dans le Pleistocene final. In D. Sonneville-Bordes, editor, La fin<br />
des temps glaciaires en Europe. Colloques Internationaux du Centre<br />
National de la Recherche Scientifique, 271: 325-332.<br />
Cassoli, PR, and A. Tagliacozzo<br />
1982. La fauna della grotta di Cala Genovesi a Levanzo. Rivista di Scienze<br />
Preistoriche, 37:124-133.<br />
Cramp, S., editor<br />
1977. Handbook of the Birds of Europe, the Middle East and North Africa;<br />
The Birds of the Western Palearctic. Volume 1,714 pages. Oxford:<br />
Oxford University Press.<br />
1983. Handbook of the Birds of Europe, the Middle East and North Africa;<br />
The Birds of the Western Palearctic. Volume 3, 913 pages. Oxford:<br />
Oxford University Press.<br />
1985. Handbook of the Birds of Europe, the Middle East and North Africa;<br />
The Birds of the Western Palearctic. Volume 4, 936 pages. Oxford:<br />
Oxford University Press.<br />
David, A.<br />
1991. Late Glacial Archaeological <strong>Res</strong>idues from Wales: A Selection. In<br />
N. Barton, A.J. Roberts, and D.A. Roe, editors, The Late Glacial in<br />
Northwest Europe. Council for British Archaeology <strong>Res</strong>earch Report,<br />
77:141-159.<br />
Del Campana, D.<br />
1946. Uccelli paleolitici degli strati basali della Grotta dei Fanciuli ai Balzi<br />
Rossi. Rivista di Scienze Preistoriche, 1:290-307.<br />
Eastham, A.<br />
1968. The Avifauna of Gorham's Cave, Gibraltar. Bulletin of the Institute<br />
of Archaeology, 1:31-42.<br />
1986. The Birds of the Cueva de Nerja. In J.F. Jorda Pardo, editor, Laprehistoria<br />
de la Cueva de Nerja (Malaga): Trabajos sobre la Cueva