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152 SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY FIGURE 11.—Area of sympatry of seabirds from Grotta Romanelli, Italy (Puffinus spp., Phalacrocorax aristotelis, Rissa tridactyla, Pinguinus impennis). (•=fossil site.) yrs. BP (Hernandez, 1995); Cala Genovesi on Levanzo Island off Sicily {Puffinus puffinus, Calonectris diomedea), which is younger than 11,180 yrs. BP (Cassoli and Tagliacozzo, 1982); and Akrotiri Aetokremnos on Cyprus {Puffinus puffinus, Phalacrocorax aristotelis), which is dated to 11,700-9000 yrs. BP (Simmons, 1991; Mourer-Chauvire, in litt.). These faunas mostly indicate conditions more or less similar to the present day, which may indicate early Holocene age. The exception is the Cueva de Nerja epipaleolithic layer (Figure 12), which indicates cool Atlantic conditions and may date largely to the cold Dryas 1 and/or Dryas 3 stadials. The fauna from Akrotiri Aetokremnos (Figure 13) also may indicate a slightly colder and more eutrophic sea around Cyprus than at present, perhaps because the fossils were deposited during the Dryas 3 stadial. UNDATED FAUNAS There are a number of faunas containing seabirds that are not at present satisfactorily dated. In these cases, consideration of the environmental information given by the seabirds may help
NUMBER 89 153 FIGURE 12.—Area of sympatry of seabirds from Cueva de Nerja, Spain (epipalaeolithic layer) (Puffinus griseus, Calonectris diomedea, Morus bassanus, Phalacrocorax aristotelis, Catharacta skua, Uria aalge, Pinguinus impennis). (•=fossil site.) in defining the age of these faunas. These faunas can be divided into two groups: faunas that indicate significantly colder conditions than at present and faunas that indicate conditions approximately similar to the present day. GROTTA PIETRO TAMPOIA, ITALY.—This is a "warm" fauna {Puffinus puffinus, Calonectris diomedea, Hydrobates pelagicus), indicating conditions similar to today, which supports Mayaud and Schaub's (1950) view that the fossils from this site are largely or wholly of Holocene age. BUCA DEL BERSAGLIERI, ITALY.—This also is a warm fauna {Puffinus puffinus, Phalacrocorax aristotelis) (Figure 13), particularly because at least some of the shearwater fossils are from the warm-water species Puffinus {puffinus) yelkouan. Very little information on the date of the deposits is available, but it seems likely that the fauna is at least partly Holocene, a hypothesis that is supported by the composition of the avifauna in general and by the presence of Rattus rattus Linnaeus (Wolf, 1938).
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NUMBER 89 153<br />
FIGURE 12.—Area of sympatry of seabirds from Cueva de Nerja, Spain (epipalaeolithic layer) (Puffinus griseus,<br />
Calonectris diomedea, Morus bassanus, Phalacrocorax aristotelis, Catharacta skua, Uria aalge, Pinguinus<br />
impennis). (•=fossil site.)<br />
in defining the age of these faunas. These faunas can be divided<br />
into two groups: faunas that indicate significantly colder<br />
conditions than at present and faunas that indicate conditions<br />
approximately similar to the present day.<br />
GROTTA PIETRO TAMPOIA, ITALY.—This is a "warm" fauna<br />
{Puffinus puffinus, Calonectris diomedea, Hydrobates pelagicus),<br />
indicating conditions similar to today, which supports<br />
Mayaud and Schaub's (1950) view that the fossils from this<br />
site are largely or wholly of Holocene age.<br />
BUCA DEL BERSAGLIERI, ITALY.—This also is a warm fauna<br />
{Puffinus puffinus, Phalacrocorax aristotelis) (Figure 13),<br />
particularly because at least some of the shearwater fossils are<br />
from the warm-water species Puffinus {puffinus) yelkouan.<br />
Very little information on the date of the deposits is available,<br />
but it seems likely that the fauna is at least partly Holocene, a<br />
hypothesis that is supported by the composition of the avifauna<br />
in general and by the presence of Rattus rattus Linnaeus<br />
(Wolf, 1938).