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152<br />

SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY<br />

FIGURE 11.—Area of sympatry of seabirds from Grotta Romanelli, Italy (Puffinus spp., Phalacrocorax aristotelis,<br />

Rissa tridactyla, Pinguinus impennis). (•=fossil site.)<br />

yrs. BP (Hernandez, 1995); Cala Genovesi on Levanzo Island<br />

off Sicily {Puffinus puffinus, Calonectris diomedea), which is<br />

younger than 11,180 yrs. BP (Cassoli and Tagliacozzo, 1982);<br />

and Akrotiri Aetokremnos on Cyprus {Puffinus puffinus, Phalacrocorax<br />

aristotelis), which is dated to 11,700-9000 yrs. BP<br />

(Simmons, 1991; Mourer-Chauvire, in litt.). These faunas<br />

mostly indicate conditions more or less similar to the present<br />

day, which may indicate early Holocene age. The exception is<br />

the Cueva de Nerja epipaleolithic layer (Figure 12), which indicates<br />

cool Atlantic conditions and may date largely to the cold<br />

Dryas 1 and/or Dryas 3 stadials. The fauna from Akrotiri<br />

Aetokremnos (Figure 13) also may indicate a slightly colder<br />

and more eutrophic sea around Cyprus than at present, perhaps<br />

because the fossils were deposited during the Dryas 3 stadial.<br />

UNDATED FAUNAS<br />

There are a number of faunas containing seabirds that are not<br />

at present satisfactorily dated. In these cases, consideration of<br />

the environmental information given by the seabirds may help

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