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NUMBER 89 147<br />

FIGURE 6.—Area of sympatry of seabirds from Arene Candide, Italy, layer P9 (Phalacrocorax aristotelis, Uria<br />

aalge, Alca torda, Fratercula arctica). (• = fossil site.)<br />

lacrocorax aristotelis (P9), Uria aalge (P4,5,7,9), Alca torda<br />

(P4,9), and Fratercula arctica (P5,9). The areas of sympatry<br />

for the faunas in layers P4, P5, and P9 are rather similar and<br />

suggest that conditions approximating those around the British<br />

Isles at the present time prevailed off the Ligurian coast during<br />

the glacial. The area of sympatry for layer P9, which has the<br />

largest number of seabird species (4), is shown in Figure 6.<br />

Conditions may have been slightly milder when layer P7 was<br />

deposited (Figure 7).<br />

The Upper Paleolithic fauna from Cueva de Nerja (UP layers)<br />

near Malaga, Spain (Figure 8), is dated to the interval<br />

16,520-13,350 yrs. BP (Hernandez, 1995) and falls within the<br />

later part of the pleniglacial, including the Lascaux interstadial<br />

and the Dryas 1 stadial. The unusually rich seabird fauna consists<br />

of Calonectris diomedea, Puffinus puffinus, P. gravis, P.<br />

griseus, Phalacrocorax aristotelis, Morus bassanus, Uria aalge,<br />

Alca torda, and Pinguinus impennis. This fauna has only a<br />

very small area of sympatry off Southwest Ireland. Such a

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