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142 SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY<br />

TABLE 2.—Composition of late Pleistocene seabird faunas. \=Fulmarus glacialis, 2=Puffinus puffinus, 3=Puffinus holei,<br />

4=Puffinus gravis, 5=Puffmus griseus, 6=Puffmus spp., 1=Calonectris diomedea, %=Hydrobates pelagicus, 9=Phalacrocorax<br />

aristotelis, 10= Morus bassanus, \\=Somateria mollissima, \2=Catharactaskua, \3=Rissa tridactyla, \4=Pagophila eburnea,<br />

\5=Alle alle, \6=Uria aalge, \l=Uria lomvia, \8=Uria sp., \9=Cepphus grylle, 20=Alca torda, 2\=Pinguinus impennis,<br />

22=Fratercula arctica.<br />

Site<br />

Cyprus<br />

Akrotiri Aetokremnos<br />

Italy<br />

Archi<br />

Arene Candide<br />

Buca del Bersaglieri<br />

Cala Genovesi a Levanzo<br />

Grotta Pietro Tampoia<br />

Grotta Romanelli<br />

Grotta dei Fanciulli<br />

Norway<br />

Blomvag<br />

Skjonghelleren<br />

Portugal<br />

Grotte de Fuminha<br />

Gruta de Figueira Brava<br />

Spain<br />

Cova den Jaume Orat<br />

Cova Nova<br />

Cueva de Nerja<br />

Devil's Tower<br />

Es Pouas<br />

Gorham's Cave<br />

Great Britain<br />

Bacon Hole<br />

Creag Nan Uamh<br />

Paviland Cave<br />

Potter's Cave<br />

Layer<br />

P9<br />

P7<br />

P5<br />

P4<br />

1. 2 t. 4<br />

1. 2 t. 3<br />

C<br />

A6<br />

EP<br />

UP<br />

UP/EP<br />

A2<br />

Bl<br />

K<br />

D-F<br />

5<br />

DATED FAUNAS<br />

2<br />

X<br />

X<br />

X<br />

X<br />

X<br />

3 4 5 6 7<br />

EEMIAN INTERGLACIAL.—Only one site can be confidently<br />

assigned to the last interglacial (Eemian/Ipswichian sensu stricto,<br />

i.e., oxygen isotope stage (IS) 5e), Bacon Hole on the Gower<br />

Peninsula in Southwest Wales. Layers D-F at this site,<br />

which have been U/Th dated to 122±9 kilo annum (Ka) BP<br />

(Stringer et al., 1986), contain a temperate avifauna, including<br />

Calonectris diomedea and Alca torda (Harrison, 1977, 1987).<br />

The current area of sympatry of these two species is situated in<br />

an area stretching from the waters southwest of the British Isles<br />

to the western Mediterranean (Figure 2). There is no sympatric<br />

breeding of these two species today, but this might be due to<br />

the absence of suitable breeding sites for seabirds between<br />

Bretagne and Northwest Spain. The occurrence of these two<br />

species in Wales supports evidence from other sources (e.g.,<br />

X<br />

X<br />

X<br />

8 9<br />

X<br />

X<br />

X<br />

10 11<br />

X<br />

12 13 14 15 16 17 18 19 20 21 22<br />

X X X<br />

X<br />

X X<br />

X X<br />

Mclntyre et al., 1972; Ruddiman and Mclntyre, 1976) that the<br />

Northeast Atlantic was somewhat warmer during the Eemian<br />

than during the present interglacial.<br />

THE EARLY WEICHSELIAN.—This is a climatically complex<br />

interval comprising two moderately cold stadials (IS 5b and IS<br />

5d) and two interstadials of nearly interglacial magnitude (ISs<br />

5a and 5c). The character of these interstadials was different in<br />

southern and northern Europe. In the south, IS 5a was, if anything,<br />

warmer than the earlier IS 5c, whereas in Scandinavia<br />

the opposite was the case (e.g., Mangerud, 1991). The extent of<br />

glaciation during the stadials is uncertain, but in Scandinavia it<br />

seems to have been mainly restricted to the mountains except<br />

in the far north. No fauna can be definitely assigned to this period,<br />

but it is possible that the fauna from Grotte de Furninha,<br />

Portugal (Tables 1, 2), for example, belongs here.<br />

X

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