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NUMBER 89<br />

the Chatham Island Pied Oystercatcher {Haematopus chathamensis),<br />

also might have become extinct.<br />

Conclusions<br />

Analysis of the abundant and well-dated fossil avian material<br />

found in sand-dune and cave deposits on several of the<br />

larger islands of the Chatham group indicates that these islands<br />

have supported a diverse, highly endemic avifauna since<br />

at least 7000 years ago. This avifauna, some 100 species in<br />

all, including many endemic land and freshwater birds, as<br />

well as a wide variety of seabirds, survived apparently unscathed<br />

until shortly after the first human arrival about 450<br />

years ago.<br />

Many birds of the Chatham Islands exemplify the evolutionary<br />

trend toward larger body size and diminished flying ability<br />

so typically found in small, isolated oceanic-island groups that,<br />

This appendix provides an annotated listing of both conventional<br />

and calibrated radiocarbon ages for samples from<br />

Chatham and Pitt islands. Samples (bone collagen or marineshell<br />

carbonate) are identified by their Rafter Radiocarbon<br />

Laboratory reference numbers (prefixed by NZA). Age data are<br />

presented as follows: conventional age based on the old (Libby)<br />

half-life of 5568 yrs (as "age ± standard deviation yrs BP");<br />

calibrated (corrected) age, given as median age ("CAL BP")<br />

where possible; and calibrated (corrected) age as a range expressed<br />

in terms of the 95% confidence interval (age ± two<br />

standard deviations). <strong>Lo</strong>cality names and grid references (GR)<br />

given for sampled sites (see Figure 1, numbered sites 1-21) are<br />

from New Zealand Topographical Map, NZMS 260, 1:50000<br />

metric series, Chatham Islands, edition 1, 1981 (Chatham Island<br />

(localities 1-17), sheet 1; Pitt Island (localities 18-21),<br />

sheet 2 ). (a.s.l.=above sea level.)<br />

CHATHAM ISLAND<br />

<strong>Lo</strong>cality 1, Waihora, Point Durham<br />

NZA 3193: Gallirallus dieffenbachii, Sutton coll. WH/VII/2<br />

Layer 1; GR 358470; Waihora Mound site, supposedly midden<br />

material; 5030 ± 68 yrs BP; ca. 5750 CAL BP;<br />

5894-5605 CAL BP.<br />

NZA 3194: Gallirallus dieffenbachii, Sutton coll. WH/VII/<br />

23 Layer 3; GR 358470; Waihora Mound site, supposedly<br />

midden material; 5237 ± 72 yrs BP; ca. 5950 CAL BP;<br />

6173-5753 CAL BP.<br />

Appendix<br />

103<br />

prior to human colonization, lacked mammalian predators.<br />

Most of the Chatham Islands land birds and waterfowl are larger<br />

than their mainland counterparts, and of the 36 prehistorically<br />

known species, at least seven were flightless and three more<br />

would have been weak fliers.<br />

The land birds of the Chatham Islands were clearly no exception<br />

to the general rule that insular species tend to be "naive"<br />

toward humans and introduced predators (Milberg and<br />

Tyrberg, 1993:229). The lethal combination of weak flight and<br />

trusting attitude predisposed them to an extraordinary vulnerability<br />

to human interference. The fossil record of the last 7000<br />

years gives no indication that any of the prehistorically known<br />

species became extinct, or even less abundant, prior to human<br />

arrival. All of the flightless and weak-flying species, and a further<br />

11 flying species, however, became extinct within a few<br />

hundred years of first human settlement through the combined<br />

effects of human perturbations.<br />

<strong>Lo</strong>cality 2, Red Bluff<br />

NZA 2610: Hemiphaga chathamica, MNZ S31065; GR<br />

469603; back-beach sequence in embayment, -800 m SSE<br />

of Te Whenuhau Trig, from uppermost stratum of coarse,<br />

yellow, shell sand (beneath windblown fine sand). Indicates<br />

minimum age for >2 m thick sequence; 985 ± 80 yrs BP;<br />

843 CAL BP; 975-689 CAL BP.<br />

<strong>Lo</strong>cality 3, <strong>Lo</strong>ng Beach, S of Henga<br />

limestone bluffs (GR 450655)<br />

NZA 1930: Fulica chathamensis, MNZ S27821; GR 465626;<br />

<strong>Lo</strong>ng Beach, in typical, stratified, consolidated, dune hummock<br />

(-4 m a.s.l.), immediately beneath eroded surface of<br />

brown sand/soil horizon (30 cm thick), now overlain by<br />

drift sand; 2560 ± 145 yrs BP; 2254 CAL BP; 2666-1924<br />

CAL BP.<br />

NZA 1931: Hemiphaga chathamensis, MNZ S27822; GR<br />

465626; <strong>Lo</strong>ng Beach, in pale brown sand at 1.5 m depth (directly<br />

underlying sample NZA 1930); 3790 ± 150 yrs BP;<br />

4109 CAL BP; 4511-3697 CAL BP.<br />

NZA 1929: Diaphorapteryx hawkinsi, MNZ S27820; GR<br />

456646; <strong>Lo</strong>ng Beach, Milton's Gully site, embedded in gray<br />

brown, hard-pan deflation surface (former interdune lake<br />

deposit?); 6660 ± 150 yrs BP; 7456 CAL BP; 7693-7207<br />

CAL BP.<br />

NZA 3246: Hemiphaga chathamensis, MNZ S33158; GR<br />

456646; <strong>Lo</strong>ng Beach, Milton's Gully site, in horizontally<br />

bedded brown sand at inland margin of and stratigraphically

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