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NUMBER 89 99<br />

FIGURE 11.—Skulls of Anatidae (dorsal views). Left to right: Anas, species undescribed (MNZ S33298, ex. H.O.<br />

Forbes collection), from "Chatham Islands"; Anas chlorotis (MNZ 14978); Tadorna variegata (MNZ 16473,<br />

male); Tadorna, species undescribed (MNZ S32830, presumed male), from Maunganui, Chatham Island. (Scale<br />

bar=30 mm.)<br />

the Te Whanga Lagoon (Turbott, 1990). The extinct species,<br />

known also from mainland New Zealand, differs in being<br />

somewhat larger, with a stouter bill and relatively shorter<br />

wings. Its bones have been found in greatest abundance at<br />

Waitangi West, at Tioriori, and at Te One Beach, near Red<br />

Bluff, in deposits ranging in age from ca. 7600 (cf. NZA 3238,<br />

locality 4) to ca. 700 (NZA 2603, locality 11) years old, but<br />

they also occur commonly in middens. The Chatham Island<br />

Duck, whose bones have been found at a considerable number<br />

of sites on Chatham Island, but not elsewhere, was robust,<br />

weighing as much as 2.5 kg (more than twice the weight of a<br />

Grey Duck), and yet with wings perhaps even a little smaller<br />

than in that species (pers. obs.). The new merganser {Mergus,<br />

species undescribed), whose bones have been found in abundance<br />

only in Te Ana a Moe Cave, was a little smaller than the<br />

recently extinct (ca. 1905), flightless Auckland Island Mergan­<br />

ser {Mergus australis; see Kear and Scarlett, 1970) and had a<br />

shorter bill and even more reduced wings (Millener, pers. obs.).<br />

EXTINCTIONS<br />

The indigenous bird faunas of remote oceanic islands, having<br />

evolved in isolation from most or all vertebrate predators,<br />

are extraordinarily vulnerable to the impact of humans (see<br />

Milberg and Tyrberg, 1993; Steadman, 1995). The land birds<br />

and waterfowl of the Chatham Islands were well adapted to<br />

their island environment. Most became larger than their mainland<br />

counterparts, many became completely or nearly flightless,<br />

some probably laid smaller clutches, and none developed<br />

or retained a fear of mammalian predators. First human settlement<br />

was almost certainly accompanied by accidental or deliberate<br />

forest clearing, and on such small islands there would

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