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NUMBER 89 93<br />

TABLE 2.—Land snails (Gastropoda: Pulmonata) identified from Te Ana a Moe Cave, Chatham Island (<strong>Lo</strong>cality<br />

16 (Figure 1)). Taxonomy follows Climo (pers. comm., 1993). Sample 1 (PRM sample 155/91) is from brown<br />

sand/soil in the uppermost 20 cm of undisturbed sediment; sample 2 (PRM sample 156/91) is from creamy white<br />

bryozoan detrital sand, at 1 m depth (-40 cm below upper surface of undisturbed sediment); sample 3 (PRM<br />

sample 120A/91) is from brown gray bryozoan detrital sand at 1.2-1.4 m depth (60-80 cm below upper surface<br />

of undisturbed sediment) in a side tunnel in S W quadrant. For each sample, abundance of each species is expressed<br />

as a percentage of sample size (n). (See also Figure 6.)<br />

Family<br />

TORNATELLINIDAE<br />

PUNCTIDAE<br />

CHAROPIDAE<br />

FLAMMULINIDAE<br />

ROTADISCIDAE<br />

CAMAENIDAE<br />

Species<br />

Lamellidea novoseelandica<br />

Serratopunctum serratocostata<br />

Litopunctum rakiura<br />

Punctum lateumbilicata<br />

Alexaoma chathamensis<br />

Dellopsis stewartensis<br />

Pryhina chathamensis<br />

Phenacharopa pseudanguicula<br />

Charopa coma<br />

Flammocharopa mayhillae<br />

Sinployea parva<br />

Huonodon hectori<br />

Mitodon wairarapa<br />

Basimocella maculata<br />

Discocharopa eta<br />

Cavellia buccinella<br />

Thalassohelix sp.<br />

TABLE 3.—Bird species identified from Te Ana a Moe Cave, Chatham Island<br />

(<strong>Lo</strong>cality 16 (Figure 1)). See "Appendix" and Table 1. (A=abundant (minimum<br />

number of individuals (MNI) >100); C=common (MNI > 10); R=rare<br />

(MNK 10); *=extinct.)<br />

Taxon<br />

MARINE SPECIES<br />

Pelecanoides urinatrix<br />

Pachyptila turtur<br />

Pachyptila crassirostris<br />

Pterodroma nigripennis<br />

Pterodroma axillaris<br />

Pterodroma magentae<br />

Pelagodroma marina<br />

Eudyptula minor<br />

Larus dominicanus<br />

TERRESTRIAL AND FRESHWATER SPECIES<br />

*Tadorna, species undescribed<br />

*Anas, species undescribed<br />

*Pachyanas chathamica<br />

*Mergus, species undescribed<br />

Falco novaeseelandiae<br />

*Gallirallus dieffenbachii<br />

*Gallirallus modestus<br />

"Diaphorapteryx hawkinsi<br />

*Fulica chathamensis<br />

*Coenocorypha chathamica<br />

Hemiphaga chathamensis<br />

*Nestor, species undescribed<br />

Cyanoramphus novaezelandiae<br />

Cyanoramphus auriceps<br />

Anthus novaeseelandiae<br />

*Bowdleria rufescens<br />

Gerygone albofrontata<br />

Rhipidura fuliginosa<br />

Petroica traversi<br />

*Anthornis melanura<br />

Prosthemadera novaeseelandiae<br />

Abundance<br />

A (at all levels)<br />

A (at all levels)<br />

R<br />

R<br />

C<br />

A (at all levels)<br />

R<br />

C<br />

R (upper level only)<br />

R (MNI=2, in upper level only)<br />

R<br />

R (MNI=1, in lower level)<br />

A (at all levels)<br />

R<br />

A (at all levels)<br />

A (at all levels)<br />

R<br />

R<br />

C<br />

R<br />

R<br />

C<br />

R<br />

R<br />

C (at all levels)<br />

R<br />

R<br />

C (at all levels)<br />

R<br />

R<br />

1 («=164)<br />

0.6<br />

-<br />

74.4<br />

-<br />

- -<br />

10.4<br />

1.2<br />

- -<br />

0.6<br />

-<br />

-<br />

5.5<br />

0.6<br />

2.4<br />

4.3<br />

Sample<br />

2(«=110)<br />

-<br />

0.9<br />

18.2<br />

4.5<br />

-<br />

-<br />

21.8<br />

-<br />

6.4<br />

-<br />

-<br />

2.7<br />

2.7<br />

35.5<br />

-<br />

8.2<br />

1.8<br />

3(«=128)<br />

1.6<br />

-<br />

40.6<br />

4.7<br />

0.6<br />

0.8<br />

13.3<br />

1.6<br />

2.3<br />

0.8<br />

1.6<br />

1.6<br />

-<br />

18.8<br />

-<br />

7.0<br />

3.9<br />

SWAMP SITES<br />

Although peat deposits and more recent swamps are widespread<br />

on Chatham Island, the conditions in them appear to<br />

have been unsuitable for the preservation of bones. Peat fires<br />

burning to considerable depths have occurred frequently on the<br />

island. Furthermore, both peats and more recent swamps seem<br />

typically to have been too acidic to allow long-term preservation<br />

of bone.<br />

Paleogeography, Ancestral Immigration, and<br />

Avifauna Change<br />

It is possible that the Chatham Islands, although isolated by<br />

a broad oceanic gap from the New Zealand mainland since the<br />

Late Cretaceous, some 80 million years ago, have provided a<br />

land mass capable of supporting viable bird populations more<br />

or less continuously for perhaps many millions of years<br />

(Fleming, 1962, 1975; Cooper and Millener, 1993). There is<br />

evidence, however, that during the late Eocene (40 Ma) and<br />

again during the Pliocene (5-2 Ma) the only emergent land in<br />

the entire Chathams group would have been a few volcanic<br />

peaks (Campbell, 1996:36). Thus colonization by the forerunners<br />

of the Holocene species may postdate the Pliocene. Further,<br />

there is some suggestion that later, in the Pleistocene,<br />

high interglacial sea-levels during the Castlecliffian (ca.<br />

1.4-0.32 Ma) may have inundated all but the highest points of<br />

Chatham Island (see Hay et al., 1970). Therefore much, if not<br />

all, of any earlier established terrestrial avifauna may have<br />

been eliminated during this period. Any ancestral avian colonists<br />

that reached the Chathams since the Cretaceous, including<br />

those still arriving today, could have done so only by fly-

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