The Madagascan Coraebina Bedel, 1921 (Coleoptera: Buprestidae ...

The Madagascan Coraebina Bedel, 1921 (Coleoptera: 

Buprestidae: Agrilinae: Agrilini): introduction and 

systematic catalogue * 

C. L. Bellamy † 

Transvaal Museum, P.O. Box 413, Pretoria, 0001 South Africa 

Bellamy, C. L., 2001. The Madagascan Coraebina Bedel, 1921 (Coleoptera: Buprestidae: Agrilinae: Agrilini): introduction 

and systematic catalogue. Annals of the Transvaal Museum 38: 57–70. 

An introduction to a vast descriptive and systematic project that will consider the large fauna of coraebine jewel 

beetles from Madagascar and the Comoro Islands is presented. The described species are summarized in a 

catalogue that contains 28 genera, one subgenus, 103 species and one subspecies. Coraebastus imperatrix 

(Obenberger), from Madagascar, is resurrected from synonymy under C. quinquepustulatus Fairmaire, from the 

Comoro Islands. Chloricala gratiosa Kerremans var. octosticta Obenberger and var. impunctata Théry and 

Pseudagrilus scintillans var. madegassus Obenberger are synonymized under their nominate species. 

Le vaste projet de description systématique de l’importante faune des Buprestes Coraebinae de Madagascar et des 

Comores est présenté. La description des espèces est résumée dans un catalogue contenant 28 genres, un 

sous-genre, 103 espèces et une sous-espèce. Coraebastus imperatrix (Obenberger), de Madagascar, est rétabli au 

rang d’espèce. Il n’est plus synonyme de C. quinquepustulatus Fairmaire des Comores. Les var. otosticta 

Obenberger et impunctata Théry du Chloricala gratiosa Kerremans ainsi que le Pseudagrillus scintillans var. 

madegassus Obenberger sont synonymisés sous le nom de leurs espèces nominales. 

Keywords/Mots Clés: Coleoptera, Buprestidae, Agrilinae, Coraebina, Madagascar. 

INTRODUCTION 

There are many accounts of a proliferation of 

species in the literature. Perhaps these come more 

frequently now with more attention being paid to 

inventorying threatened habitats as world populations 

rise and development invades once pristine 

tracts of nature. In my experience, there is nothing 

that compares to the vast and largely undescribed 

jewel beetle subtribe Coraebina Bedel (1921) 

(Coleoptera: Buprestidae: Agrilinae: Agrilini) from 

Madagascar. This is the second contribution in a 

series that is anticipated to take many years to 

complete; the first was by Bellamy (1997). 

During the latter half of the 20th century, French 

and Malagasy entomologists discovered that 

nest-provisioning wasps (Hymenoptera: Sphecoidea: 

Philanthidae: Cerceris Latreille) were using 

small jewel beetles as their primary prey. The two 

principal collectors on the majority of specimen 

data labels are P. Vadon and A. Peyrieras. The beetles 

are paralyzed by the sting of female wasps and 

then carried back to the nests, whereupon an egg is 

laid and the emerging larva feeds on the beetle 

before pupating inside the hollow body. By following 

the wasps and learning to recognize their burrows 

*Third contribution to the knowledge of Madagascar Buprestidae. 

† 

Present address: Entomology Section, Natural History Museum of 

Los Angeles County, 900 Exposition Blvd., Los Angeles, California, 

90007, U.S.A. E-mail: cbellamy@nhm.org 

or nests, these entomologists were able to dig up 

many comatose beetles, which lay in perfect or 

near-perfect condition, and return them for preparation 

and permanent museum storage. 

The predator–prey relationship between certain 

species of Cerceris and buprestids has been widely 

reported for various parts of the world (e.g. in general: 

Cartwright, 1931; Inglebert, 1993; Scullen and 

Wold, 1969; Trinidad: Fisher, 1944; Callan, 1990; 

North America: Grossbeck, 1912; Linsley and 

MacSwain, 1956; Evans and Rubinik, 1978; 

Kurczewski and Miller, 1984; Hook and Evans, 

1991; Japan: Ohmomo and Usuba, 1997). 

Several colleagues (W. F. Barr, H. A. Hespenheide, 

S. G. Wellso, pers. comm.) had told me about the 

collection of Madagascan buprestids seen in the 

Museum National d’Histoire Naturelle, Paris, during 

their respective research visits in the 1970s and 

1980s, but until I experienced this firsthand during 

my initial visit to the museum in late 1995, I really 

could not understand the breadth of this collection. 

There are thousands of specimens, prepared and 

labelled, representing a large number of mostly 

undescribed species. From an initial attempt to 

sort and arrange these specimens by the former 

head of the Coleoptera Department, the late 

A. Descarpentries, an estimate of the size of this 

fauna is between 500–600 undescribed species, 

which may require 20–30 new genera to contain

58 ANNALS OF THE TRANSVAAL MUSEUM, VOLUME 38, 2001 

them. It seems clear from the literature, as well as 

from the examples of other buprestid prey in this 

material, that the Cerceris predators are not 

particular to coraebine buprestids, nor possibly 

even just buprestids. There are large series of other 

buprestid species (e.g., Sponsor Gory and Laporte, 

see Descarpentries, 1965; Anthaxia Eschscholtz, 

see Descarpentries, 1967) that are each of about 

the same general size and shape as the many 

coraebine species and this suggests that the 

predators are perhaps more selective about the 

physical characteristics of the prey, perhaps to be 

able to contain them within specific morphological 

adaptations to maintain a hold. Krombein (1981: 8) 

correlated structure and function when he described 

and figured the ‘buprestid clamp’ found in 

members of the ‘bupresticida group’ of Cerceris, 

including species from Sri Lanka, Europe and North 

America. 

The present paper is intended as an introduction 

to the many nomenclatural acts and descriptive 

papers that must follow. By introducing the subject 

and providing an accounting of the current systematics 

of the known Madagascan coraebine fauna, all 

subsequent efforts can focus on the description of 

new taxa and several of the large genera which have 

never been revised, e.g. Entomogaster Saunders. 

The catalogue included herein is presented in full, 

indicating all taxa and the nomenclatural history for 

each species. This effort to present a full catalogue 

of the previously described taxa of this subtribe is 

undertaken now to bring together all of the available 

information on a large group of species that have 

mostly appeared in random and disparate descriptive 

publications and earlier more general catalogues. 

The chronology of catalogues that have 

treated these taxa, in part, are: Gemminger and Harold, 

1869; Saunders, 1871; Kerremans, 1885, 1892; 

Alluaud, 1900; Kerremans, 1903; Théry, 1905; Marie 

and Lesne, 1917; Obenberger, 1934; and Bellamy, 

1985. The work of Obenberger (1934) was the fourth 

of six fascicles of the two volumes on Buprestidae in 

the Coleopterorum Catalogue, the last effort to catalogue 

the world’s beetle fauna. Many of the taxa of 

Madagascan Coraebina were described subsequent 

to that catalogue and have not been found in 

the literature after their respective original descriptions. 

For future geographical notes, distributional 

accounts and attempts to summarize the range of 

particular taxa within the Madagascan habitats, I 

note here that a great many of the older taxa of this 

group were described from a type locality no more 

precise than ‘Madagascar’. However, considering 

the wealth of specimens of many of these taxa that 

were harvested, some detail will be filled in as 

species are described and genera are revised. As a 

reference for Madagascan place names, the work 

of Viette (1991) will be the standard. 

The described complement of these beetles is 28 

genera, one subgenus, 103 species and one subspecies. 

The single related taxon, Coraebastus 

quinquepustulatus Fairmaire, from the Comoro 

Islands is included with the reinstatement from 

synonymy of Coraebastus imperatrix (Obenberger) 

from Madagascar. In the catalogue, the genera and 

species are listed alphabetically. With so much 

descriptive work to be done, phylogenetic predictions 

are premature, since each subsequent 

descriptive paper will alter the conclusions about 

transformation character state series and 

synapomorphies. At the end of the catalogue are 

listed 11 species names that were originally 

described in one of two older coraebine genera. 

These are listed as ‘undefined placement’ pending 

better understanding of relationships with the many 

new species amongst which some of these will 

undoubtedly fit. Following current trends in dealing 

with infrasubspecific names, I will propose all 

varietal names listed below as new synonyms (see 

Bellamy & Peterson, 2000). 

Abbreviations used herein are: nom. nov. = 

nomen novum; nom. nud. = nomen nudum; nom. 

superfl. = superfluous name, i.e. a replacement 

name that was unnecessary; orig. desc. = original 

description; rev. = revision; spp. key = species key; 

syn. = synonym; syn. nov. = new synonym; 

unvn. = unavailable name, i.e. infrasubspecific 

names, nomina nuda; var. = variety, variation. 

SYSTEMATIC CATALOGUE OF THE 

COROEBINA OF THE MADAGASCAN 

REGION 

Genus 1. ANGATRA Descarpentries 

Angatra Descarpentries, 1969: 599–600 (orig. 

desc.); Bellamy, 1985: 424. 

Type species: Angatra magna Descarpentries 

(original designation). 

REMARKS. This genus is endemic to Madagascar 

and apparently has no close relationships to the 

Afrotropical fauna. 

1. magna Descarpentries 

Angatra magna Descarpentries, 1969: 600–601 

(orig. desc.), fig. 1. 

2. parva Descarpentries 

Angatra parva Descarpentries, 1969: 601–602 


Genus 2. ANTANAMBIA Descarpentries 

Antanambia Descarpentries, 1975: 19–20 (orig.

MADAGASCAN CORAEBINA (COLEOPTERA): SYSTEMATIC CATALOGUE 59 

desc.); Bellamy, 1985: 425. 

Type species: Antanambia nigricans 

Descarpentries (original designation). 


and it is not particularly close to any coraebine 

lineage in sub-Saharan Africa. 

3. nigricans Descarpentries 

Antanambia nigricans Descarpentries, 1975: 

21–22 (orig. desc.), figs 1, 3–5. 

Genus 3. BRACHYDORA Obenberger 

Brachydora Obenberger, 1923: 30–31 (orig. 

desc.); 1934: 794; Bellamy, 1985: 423. 

Type species: Brachydora monstrum 

Obenberger (original designation). 


and is quite distinct from the Afrotropical fauna. 

4. crassa Théry 

Brachydora crassa Théry, 1937: 87–89 (orig. 

desc.), fig. 3. 

5. deformis (Fairmaire) 

Discoderes deformis Fairmaire, 1901: 163 (orig. 

desc.); Kerremans, 1903: 223; Marie and 

Lesne, 1917: 40. 

Paradora deformis: Théry, 1905: 125. 

Brachydora deformis: Obenberger, 1923: 32; 

1934: 795. 

6. granulum (Fairmaire) 

Discoderes granulum Fairmaire, 1902b: 

350–351 (orig. desc.); Théry, 1905: 116; 

Marie and Lesne, 1917: 40 

Brachydora granulum: Obenberger, 1928: 342; 

1934: 795. 

7. monstrum Obenberger 

Brachydora monstrum Obenberger, 1923: 31–32 

(orig. desc.); 1934: 795. 

8. sicardi (Théry) 

Paradora sicardi Théry, 1912: 118–119 (orig. 

desc.,) 

Brachydora sicardi: Obenberger, 1923: 31; 1934: 

795. 

Genus 4. CHLORICALA Kerremans 

Chloricala Kerremans, 1893: 119–120 (orig. 

desc.); Théry, 1905: 120; Bellamy, 1985: 424. 

Type species: Chloricala gratiosa Kerremans 

(original monotypy). 

Chloricalla (lapsus calami): Kerremans, 1900: 

319; 1903: 231. 


and shows no close similarities to the Afrotropical 

fauna. The two varietal names deserve no recognition 

and are technically invalid (ICZN 1999, Article 

45.5, 45.6). 

9. balachowskyi Descarpentries 

Chloricala balachowskyi Descarpentries, 1974: 

582–584 (orig. desc.), figs 7, 8. 

10. gratiosa Kerremans 

Chloricala gratiosa Kerremans, 1893: 120 (orig. 

desc.); 1900: 319; 1903: 231, plate 2, fig. 11; 

Théry, 1905: 120; Obenberger, 1934: 813; 

1935b: 20; Théry, 1936a: 59. 

Chloricala gratiosa var. octosticta Obenberger, 

1935b: 20 (orig. desc.); 1934: 813; Théry, 

1936a: 59. Syn. nov. 

Chloricala gratiosa var. impunctata Théry, 1936a: 

60–61 (orig. desc.). Syn. nov. 

Genus 5. COMPSOGLYPTA Fairmaire 

Compsoglypta Fairmaire, 1904: 233 (orig. 

desc.); Kerremans, 1910: 3 (syn. Psiloptera), 

347; Marie and Lesne, 1917: 30; Obenberger, 

1928: 339; 1934: 795; Bellamy, 1985: 423. 

Type species: Compsoglypta perrieri Fairmaire 


Compsoglypha (lapsus calami): Théry, 1905: 

177; Obenberger, 1934: 75; Bellamy, 1985: 

423. 


and its placement in Psilopterina or Coraebina has 

been somewhat confused as indicated by the 

synonymy above. I have examined the types of the 

Fairmaire taxon and am confident that this is a 

member of the coraebine lineage, albeit very divergent. 

11. perrieri Fairmaire 

Compsoglypta perrieri Fairmaire, 1904: 233–234 

(orig. desc.); Marie and Lesne, 1917: 30; 

Obenberger, 1926: 197; 1928: 339; 1934: 

795. 

Compsoglypha perrieri: Théry, 1905: 177. 

Psiloptera sculpticollis Kerremans, 1911: 347 

(orig. desc., nom. superfl. perrieri Fairmaire); 

Obenberger, 1926: 197; 1928: 339; 1934: 

795. 

Genus 6. CORAEBASTUS Fairmaire 

Coraebastus Fairmaire, 1896: 456–457 (orig. 

desc.); Kerremans, 1903: 223; Théry, 1905: 

116; Marie and Lesne, 1917: 31; Obenberger, 

1934: 809; Descarpentries, 1978: 107; 

Bellamy, 1985: 424.


Type species: Coraebastus quinquepustulatus 

Fairmaire (original monotypy). 

Annietta Obenberger, 1931: 184; 1934: 809; 

Descarpentries, 1978: 108 (synonym of 

Coraebastus); Bellamy, 1985: 424. 

Type species: Annietta imperatrix Obenberger 



and the Comoro Islands. While I am in agreement 

with Descarpentries (1978) about the generic 

synonymy, I do not agree with the species synonymy. 

I have examined at least one specimen of each 

species from Madagascar and the Comores and 

find species level distinctions that validate both 

taxa. This is not surprising given the different 

species in other buprestid genera found in 

these two places (e.g. Polybothris Spinola, Chalcophoropsis 

monochroma from the Comores 

(Gianasso 1999)). 

12. imperatrix (Obenberger), name reinstated 

Annietta imperatrix Obenberger, 1931: 184–185 

(orig. desc.); 1934: 809. 

Coraebastus imperatrix: Descarpentries, 1978: 

108 (syn. quinquepustulatus). 

13. quinquepustulatus Fairmaire 

Coraebastus quinquepustulatus Fairmaire, 

1896: 456 (orig. desc.); Alluaud, 1900: 197; 

Kerremans, 1903: 223; Théry, 1905: 116; 

Marie and Lesne, 1917: 31; Obenberger, 

1934: 809; Descarpentries, 1978: 107. 

Genus 7. CORAEBUS Gory and Laporte 

Coraebus Gory and Laporte, 1839: 1–2 (orig. 

desc.); Gemminger and Harold, 1869: 1429; 

Saunders, 1871: 104; Kerremans, 1892: 227; 

1893: 117; Théry, 1905: 122; Obenberger, 

1934: 817; Bellamy, 1985: 425; Holynski, 

1993: 15. 

Type species: Buprestis undatus Fabricius (subsequent 

designation: Descarpentries and 

Villiers, 1967: 471). 

REMARKS. This genus is widespread in the 

Palaearctic and Oriental Regions. Representation in 

the Afrotropical fauna was eliminated by the generic 

transfers of Bellamy (1988) and the placement of 

C. insignis in Coraebus is suspect. 

14. insignis Kerremans 

Coraebus insignis Kerremans, 1898: 83 (orig. 

desc.); Alluaud, 1900: 197; Kerremans, 1903: 

233; Théry, 1905: 122; Obenberger, 1934: 

830. 

Genus 8. DISCODEROPSIS Théry 

Discoderopsis Théry, 1930b: 168–169 (orig. 

desc.); Obenberger, 1934: 795; Bellamy, 

1985: 423. 

Type species: Discoderopsis incurvata Théry 



and is putatively close to the Afrotropical lineage 

which contains Pseudokerremansia and Cupriscobina, 

both Bellamy and Holm (1985). 

15. nigrovirens (Fairmaire) 

Discoderes nigrovirens Fairmaire, 1899: 

116–117 (orig. desc.); Alluaud, 1900: 197; 


Marie and Lesne, 1917: 40. 

Phlocteis nigrovirens: Obenberger, 1928: 341; 

1934: 798. 

Discoderopsis nigrovirens: Théry, 1936a: 60. 

Discoderopsis incurvatus Théry, 1930b: 

169–170 (orig. desc.); Obenberger, 1934: 

795; Théry, 1936a: 60 (synonym of 

nigrovirens). 

Genus 9. ENTOMOGASTER Saunders 

Entomogaster Saunders, 1871: 102; Kerremans, 

1892: 223; 1893: 122; Fairmaire, 1899: 

114–115 (description); Kerremans, 1900: 

319; 1903: 225; Théry, 1905: 117; 

Obenberger, 1928: 340; Théry, 1930b: 168; 

Obenberger, 1934: 808; Théry, 1943: 149; 

Bellamy, 1985: 424; Holynski, 1993: 15; 

Bellamy, 1997: 78. 

Type species: Anthaxia sexpunctatus Gory and 

Laporte (original monotypy). 

Entomoderes Fairmaire, 1903a: 30 (in part, 

lapsus calami for Entomogaster); Marie and 

Lesne, 1917: 45. 


and shows no particular similarities to any lineage of 

the Afrotropical fauna. There are many hundreds of 

specimens of Entomogaster in the study material in 

Paris. 

16. aequalipennis Obenberger 

Entomogaster aequalipennis Obenberger, 1931: 

197–198 (orig. desc.); 1934: 808. 

17. amplicollis Fairmaire 

Entomogaster amplicollis Fairmaire, 1900: 

243–244 (orig. desc.); Kerremans, 1900: 319; 

1903: 226; Théry, 1905: 118; Marie and 

Lesne, 1917: 46; Obenberger, 1928: 340; 

1934: 808.


18. apollo Obenberger 

Entomogaster apollo Obenberger, 1946: 

160–161 (orig. desc.). 

19. asperulus Théry 

Entomogaster asperulus Théry, 1905: 120, 179 

(orig. desc.), fig. 20; Obenberger, 1928: 340; 

1934: 808. 

20. carli Théry 

Entomogaster carli Théry, 1936b: 697–699 (orig. 


21. descarpentriesi Théry 

Entomogaster descarpentriesi Théry, 1937: 

95–96 (orig. desc.), fig. 6. 

22. funebris Théry 

Entomogaster funebris Théry, 1937: 98–100 


23. grandis Théry 

Entomogaster grandis Théry, 1937: 100–102 


24. griseofasciatus Obenberger 

Entomogaster griseofasciatus Obenberger, 

1946: 162–163 (orig. desc.). 

25. hoschecki Théry 

Entomogaster hoschecki Théry, 1937: 96–98 


26. iris Obenberger 

Entomogaster iris Obenberger, 1946: 160 (orig. 

desc.). 

27. kerremansi Fairmaire 

Entomogaster kerremansi Fairmaire, 1899: 115 

(orig. desc.); Alluaud, 1900: 197; Kerremans, 

1900: 319; 1903: 225; Théry, 1905: 118; Marie 

and Lesne, 1917: 46; Obenberger, 1928: 340; 

1934: 808; Bellamy, 1997: 78. 

Entomogaster lantzi Théry (nom. nud., teste 

Obenberger, 1928: 340); Obenberger, 1934: 

808. 

28. maculipictus Obenberger 

Entomogaster maculipictus Obenberger, 1931: 

197 (orig. desc.); 1934: 808. 

29. mandrakanus Obenberger 

Entomogaster mandrakanus Obenberger, 1946: 

161–162 (orig. desc.). 

30. mocquerysi Théry 

Entomogaster mocquerysi Théry, 1905: 120 

(orig. desc.); Obenberger, 1928: 340; 1934: 

808. 

31. modestus Fairmaire 

Entomogaster modestus Fairmaire, 1899: 

115–116 (orig. desc.); Alluaud, 1900: 198; 

Kerremans, 1900: 319; 1903: 226; Théry, 

1905: 118; Marie and Lesne, 1917: 46; 

Obenberger, 1928: 340; 1934: 808; Bellamy, 

1997: 78. 

32. octoguttatus (Kerremans) 

Agrilus octoguttatus Kerremans, 1894: 353 (orig. 

desc.). 

Entomogaster octoguttatus: Fairmaire, 1899: 

116; Alluaud, 1900: 198; Kerremans, 1900: 

319; 1903: 225; Théry, 1905: 118; 

Obenberger, 1928: 340; 1934: 809. 

33. posticalis (Fairmaire) 

Entomoderes posticalis Fairmaire, 1903a: 29 

(orig. desc.). 

Entomogaster posticalis: Fairmaire, 1903b: 201– 

202; Théry, 1905: 119; Marie and Lesne, 1917: 

45, 46; Obenberger, 1928: 340; 1934: 809. 

34. pumilio Obenberger 

Entomogaster pumilio Obenberger, 1946: 

163–164 (orig. desc.). 

35. rufobasalis (Fairmaire) 

Cisseis rufobasalis Fairmaire, 1897: 380 (orig. 

desc.); Alluaud, 1900: 197. 

Entomogaster rufobasalis: Kerremans, 1903: 


340; 1934: 809. 

36. sexpunctatus (Gory and Laporte) 

Anthaxia sexpunctatus Gory and Laporte, 1839: 

36–37 (orig. desc.); Gemminger and Harold, 

1869: 1391. 

Entomogaster sexpunctatus: Saunders, 1871: 

102; Kerremans, 1892: 223; 1893: 122; 

Alluaud, 1900: 198; Kerremans, 1900: 319; 


1928: 340; 1934: 809; Bellamy, 1997: 78. 

Genus 10. EUMORPHOCERUS Théry 

Eumorphocerus Théry, 1930b: 166–167 (orig. 

desc.); Obenberger, 1934: 809; 

Descarpentries, 1968: 934–935 (spp. key); 

Bellamy, 1985: 424. 

Type species: Eumorphocerus laticornis Théry 



and is quite distinct from the Afrotropical fauna.


37. costipennis Descarpentries 

Eumorphocerus costipennis Descarpentries, 

1968: 935 (orig. desc.). 

38. laticornis Théry 

Eumorphocerus laticornis Théry, 1930b: 

167–168 (orig. desc.); Obenberger, 1934: 

809; Descarpentries, 1968: 934. 

39. peyrierasi Descarpentries 

Eumorphocerus peyrierasi Descarpentries, 

1968: 935 (orig. desc.), fig. 15. 

40. vadoni Descarpentries 

Eumorphocerus vadoni Descarpentries, 1968: 

935 (orig. desc.), fig. 14. 

Genus 11. EVIMANTIUS Kerremans 

Evimantius Kerremans, 1891: clxiii (orig. desc.); 

1892: 237; 1893: 118; 1903: 245; Théry, 1905: 

123; 1930b: 168; Obenberger, 1934: 810; 

Bellamy, 1985: 424; Holynski, 1993: 15; 

Bellamy, 1997: 77. 

Type species: Evimantius rufopictus Kerremans 

(subsequent designation: Bellamy 1997). 

Evimantius Deyrolle, 1864: 113 (nom. nud.); 

Saunders, 1871: 137. 

Type species: Evimantius rufopictus La Férte 



and seems close to species of Anaphlocteis 

Bellamy in the Afrotropical fauna. 

41. curvicollis Kerremans 

Evimantius curvicollis Kerremans, 1891: clxiv 

(orig. desc.); 1892: 237; Alluaud, 1900: 198; 


Obenberger, 1934: 810; Bellamy, 1997: 77. 

42. rufopictus Kerremans 

Evimantius rufopictus Kerremans, 1891: clxiv 

(orig. desc.); 1892: 237; Alluaud, 1900: 198; 

Kerremans, 1903: 245, plate 3, fig. 4; Théry, 

1905: 123; Obenberger, 1934: 810; Bellamy, 

1997: 77. 

Evimantius rufopictus La Férte (nom. nud.) vide 

Deyrolle, 1864: 113. 

Evimantius olsufiewi Hoscheck, 1931: 733–735 

(orig. desc.); Obenberger, 1934: 810; 

Bellamy, 1997: 77 (syn. rufopictus). 

Evimantius strandi Obenberger, 1931: 181 (orig. 

desc.); 1934: 810; Bellamy, 1997: 77 (syn. 

rufopictus). 

Evimantius embriki Obenberger, 1936: 139 (orig. 

desc.); Bellamy, 1997: 77 (syn. rufopictus). 

Genus 12. GIGANTOCORAEBUS Obenberger 

Gigantocoraebus Obenberger, 1942: 12–13 

(orig. desc.); Bellamy, 1985: 425. 

Type species: Buprestis lumbaris Klug (original 

designation). 

REMARKS. This genus is endemic to Madagascar, 

quite distinct from other members of the Madagascan 

and Afrotropical faunas and is not represented 

in the Cerceris-collected material. 

43. lumbaris (Klug) 

Buprestis lumbaris Klug, 1833: 62 (orig. desc.). 

Coraebus lumbaris: Gemminger and Harold, 

1869: 1431. 

Discoderes lumbaris: Saunders, 1871: 109; 

Kerremans, 1892: 236; Alluaud, 1900: 197. 

Phlocteis lumbaris: Kerremans, 1903: 257; 

Théry, 1905: 125; Obenberger, 1934: 798. 

Gigantocoraebus lumbaris: Obenberger, 1942: 

13, fig. 5. 

Genus 13. JAROSLAVIA Obenberger 

Jaroslavia Obenberger, 1942: 11–12 (orig. 

desc.); Bellamy, 1985: 424. 

Type species: Jaroslavia ianthinipennis 


REMARKS. This monotypic genus is endemic to 

Madagascar. 

44. ianthinipennis Obenberger 

Jaroslavia ianthinipennis Obenberger, 1942: 12 


Genus 14. MANDRITSARIA Obenberger 

Mandritsaria Obenberger, 1942: 7–8 (orig. 

desc.); Descarpentries, 1968: 932–933 (spp. 

key); Bellamy, 1985: 425. 

Type species: Amorphosoma inaequale Gory 

and Laporte (original designation). 


and shows no particular affinity to any group of the 

Afrotropical fauna. 

45. ambositrensis Descarpentries 

Mandritsaria ambositrensis Descarpentries, 

1968: 933 (orig. desc.). 

46. antamponensis Descarpentries 

Mandritsaria antamponensis Descarpentries, 

1968: 932 (orig. desc.). 

47. catalai Descarpentries 

Mandritsaria catalai Descarpentries, 1968: 933 

(orig. desc.), fig. 13.


48. inaequalis (Gory and Laporte) 

Amorphosoma inaequale Gory and Laporte, 

1839: 11–12 (orig. desc.). 

Discoderes inaequalis: Gemminger and Harold, 

1869: 1433; Saunders, 1871: 109; Kerremans, 

1892: 236; 1903: 223; Alluaud, 1900: 

196; Théry, 1905: 114. 

Phlocteis inaequalis: Obenberger, 1934: 798. 

Mandritsaria inaequalis: Obenberger, 1942: 8, 

fig. 4; Descarpentries, 1968: 933. 


Mandritsaria peyrierasi Descarpentries, 1968: 

933 (orig. desc.). 

50. sicardi Descarpentries 

Mandritsaria sicardi Descarpentries, 1968: 933 



Mandritsaria vadoni Descarpentries, 1968: 932 


Genus 15. MAROANTSETRA Théry 

Maroantsetra Théry, 1937: 102–103 (orig. desc.); 

Bellamy, 1985: 425. 

Type species: Maroantsetra superba Théry (original 

monotypy). 


and is quite distinct from the Afrotropical fauna. 

Several new species await description in the Paris 

museum. 

52. superba Théry 

Maroantsetra superba Théry, 1937: 103–104 


Genus 16. MELIBOEUS Deyrolle 

Meliboeus Deyrolle, 1864: 132 (orig. desc.); 

Gemminger and Harold, 1869: 1429; 

Saunders, 1871: 107; Obenberger, 1934: 

880; Bellamy, 1985: 426. 

Type species: Agrilus episcopalis Mannerheim 

(subsequent designation: Schaefer 1949: 

333). 

REMARKS. This very speciose genus is widespread 

in the Old World except for Australasia. 

53. albomaculatus Théry 

Meliboeus albomaculatus Théry, 1937: 108–110 


54. bipustulatus (Fairmaire) 

Entomogaster bipustulatus Fairmaire, 1901: 161 

(orig. desc.); Kerremans, 1903: 226; Théry, 

1905: 118; Marie and Lesne, 1917: 46; Théry, 

1930b: 168. 

Meliboeus bipustulatus: Obenberger, 1934: 808, 

899. 

55. cyanipennis (Fairmaire) 

Entomogaster cyanipennis Fairmaire, 1903b: 

201 (orig. desc.); Théry, 1905: 119; Marie and 

Lesne, 1917: 46; Théry, 1930b: 168; 

Obenberger, 1928: 340. 

Meliboeus cyanipennis: Obenberger, 1934: 808, 

901. 

56. hovanus Théry 

Meliboeus hovanus Théry, 1937: 107 (orig. 


57. insipidus Théry 

Meliboeus insipidus Théry, 1905: 123 (orig. 

desc.), fig. 21; Obenberger, 1934: 902. 

58. miliaris (Gory and Laporte) 

Agrilus miliaris Gory and Laporte, 1839: 32 (orig. 

desc.); Gemminger and Harold, 1869: 1442. 

Meliboeus miliaris: Saunders, 1871: 105; 

Kerremans, 1892: 232; Alluaud, 1900: 198; 


Obenberger, 1934: 903. 

59. vagecostatus Théry 

Meliboeus vagecostatus Théry, 1937: 108 (orig. 

desc.), fig.13. 

Genus 17. MIDONGYA Obenberger 

Midongya Obenberger, 1942: 9–10 (orig. desc.); 

Bellamy, 1985: 425. 

Type species: Midongya mirabilis Obenberger 



Madagascar and at least one additional species is 

present in the Paris collection. 

60. mirabilis Obenberger 

Midongya mirabilis Obenberger, 1942: 10–11 


Genus 18. PARADORA Kerremans 

Paradora Kerremans, 1900: 323 (orig. desc.); 


1922: 127; 1934: 796 (syn. Phlocteis); 1935a: 

87; Bellamy, 1985: 424; 1986: 128; 1991: 5; 

1997: 76. 

Type species: Buprestis pumicatus Klug (subsequent 

designation: Bellamy, 1997). 

REMARKS. This genus is endemic to Madagascar


and is quite distinct from the Afrotropical fauna 

despite any relationship implied by the name 

Paradorella Obenberger from South Africa (see 

Bellamy, 1989). 

Subgenus Paradora (s. str.) 

61. impetigiosus (Gory) 

Coraebus impetigiosus Gory, 1841: 277–278 

(orig. desc.), plate 46, fig. 269. 

Discoderes impetigiosus: Kerremans, 1892: 236. 

Paradora impetigiosus: Kerremans, 1900: 323; 

1903: 248; Obenberger, 1935a: 87; Bellamy, 

1997: 76. 

62. pumicatus (Klug) 

Buprestis pumicatus Klug, 1833: 62 (orig. desc.). 

Coroebus pumicatus: Gory and Laporte, 1839: 

6; Gemminger and Harold, 1869: 1431. 

Discoderes pumicatus: Saunders, 1871: 109; 

Kerremans, 1892: 237; Alluaud, 1900: 197. 

Paradora pumicatus: Kerremans, 1900: 323; 


1934: 799; 1935a: 87; Bellamy, 1997: 76. 

63. purpureosignata Hoschek 

Paradora purpureosignata Hoschek, 1931: 

735–736 (orig. desc.). 

Phlocteis purpureosignata: Obenberger, 1934: 

799. 

64. saxosicollis (Fairmaire) 

Discoderes saxosicollis Fairmaire, 1897: 381 

(orig. desc.); Alluaud, 1900: 197. 

Paradora saxosicollis: Kerremans, 1900: 323; 

1903: 249; Théry, 1905: 124; Bellamy, 1997: 

76. 

Discoderes saxosicollis: Marie and Lesne, 1917: 

40. 

Phlocteis saxosicollis: Obenberger, 1928: 341; 

1934: 799; 1935a: 87. 

65. zonata (Kerremans) 

Phlocteis zonata Kerremans, 1899: 277–278 

(orig. desc.); Alluaud, 1900: 198. 

Paradora zonata: Kerremans, 1900: 323; 1903: 


799; 1935a: 87; Bellamy, 1997: 76. 

Subgenus Speophora Théry 

Paradora (Speophora) Théry, 1937: 91–92 (orig. 

desc.); Bellamy, 1985: 424; 1997: 77. 

Type species: Paradora (Speophora) seyrigi 

Théry (original monotypy). 

66. corinthia (Fairmaire) 

Discoderes corinthia Fairmaire, 1901: 162 (orig. 


115; Marie and Lesne, 1917: 40. 

Phlocteis corynthius Obenberger, 1928: 341 

(lapsus calami); Bellamy,1997: 77. 

Phlocteis corynthia Obenberger, 1934: 796 

(lapsus calami); Bellamy, 1997: 77. 

Paradora (Speophora) corinthia: Bellamy, 1997: 

77. 

Paradora (Speophora) seyrigi Théry, 1937: 

92–94 (orig. desc.), fig. 5; Bellamy 1997: 77 

(syn. corinthia). 

Genus 19. PARANASTELLA Obenberger 

Paranastella Obenberger, 1931: 198 (orig. 

desc.); 1934: 808; Descarpentries, 1984: 57; 

Bellamy, 1985: 424 

Type species: Paranastella strandi Obenberger 



and despite the generic name does not seem 

particularly close to Nastella Kerremans from South 

Africa (see Bellamy, 1989). A third species has been 

located and will be described in the near future. 

67. strandi Obenberger 

Paranastella strandi Obenberger, 1931: 198–199 

(orig. desc.); 1934: 808; Descarpentries, 

1984: 58, fig. 3. 

68. villiersi Descarpentries 

Paranastella villiersi Descarpentries, 1984: 

57–58 (orig. desc.), figs 1, 2. 

Genus 20. PEYRIERASINA Descarpentries 

Peyrierasina Descarpentries, 1975: 22–24 (orig. 

desc.); Bellamy, 1985: 424. 

Type species: Peyrierasina complanata 

Descarpentries (original designation). 


Madagascar and quite distinct from its tribal 

relatives. 

69. complanata Descarpentries 

Peyrierasina complanata Descarpentries, 1975: 

24–25 (orig. desc.), figs 2, 6–8. 

Genus 21. PHLOCTEIS Kerremans 

Phlocteis Kerremans, 1893: 118 (orig. desc.); 

1903: 257; Théry, 1905: 10, 125; Obenberger, 

1928: 341; 1934: 795; Bellamy, 1985: 423; 

1986: 128 (rev., spp. key); 1988: 112; 1991: 5. 

Type species: Buprestis exasperata Swartz (original 

designation). 

REMARKS. This genus is restricted to the Afro-


tropical region and Madagascar (see Bellamy, 

1986). 

70. hova Théry 

Phlocteis hova Théry, 1937: 89–91 (orig. desc.), 

fig. 4. 

71. humeralis (Waterhouse) 

Discoderes humeralis Waterhouse, 1887: 293 

(orig. desc.); Kerremans 1892: 236; Alluaud, 

1900: 196. 

Phlocteis humeralis: Kerremans, 1903: 257; 

Théry, 1905: 125; Obenberger, 1928: 341; 

1934: 797. 

Genus 22. PSEUDAGRILUS Laporte 

Pseudagrilus Laporte, 1835: 165 (orig. desc.); 

Gory and Laporte, 1839: 1; Deyrolle, 1864: 

116; Gemminger and Harold, 1869: 1434; 


1893: 119; Théry, 1930a: 122 (rev., spp. key); 

Obenberger, 1934: 872; Bellamy, 1985: 426; 

1988: 111; 1991: 8; Holynski, 1993: 15. 

Type species: Pseudagrilus splendidus Laporte 


Entomoderes Fairmaire, 1903a: 30 (in part, 

lapsus calami for Entomogaster); Marie and 

Lesne, 1917: 45; Obenberger, 1934: 874; 

Théry, 1955: 397. 

REMARKS. This genus is restricted to the 

Afrotropical region and Madagascar. 

72. scintillans (Fairmaire) 

Entomoderes scintillans Fairmaire, 1903a: 30 


Entomogaster scintillans: Fairmaire, 1903b: 201; 

Théry, 1905: 119; Marie and Lesne, 1917: 45, 

46. 

Pseudagrilus scintillans: Obenberger, 1934: 874. 

Pseudagrilus scintillans var. madegassus 

Obenberger, 1924: 112; 1934: 874 (var. 

scintillans). Syn. nov. 

Genus 23. PSEUDOCORAEBUS Théry 

Pseudocoraebus Théry, 1905: 121 (orig. desc.); 

Obenberger, 1934: 813; Bellamy, 1985: 424. 

Type species: Pseudocoraebus mocquerysi 

Théry (original monotypy). 


and is distinct from the Afrotropical fauna. 

73. mocquerysi Théry 

Pseudocoraebus mocquerysi Théry, 1905: 

121–122 (orig. desc.), fig. 18; Obenberger, 

1934: 813. 

74. strandi Obenberger 

Pseudocoraebus strandi Obenberger, 1931: 183 

(orig. desc.); 1934: 813. 

Genus 24. SAMBIRANIA Obenberger 

Sambirania Obenberger, 1942: 8 (orig. desc.); 


Bellamy, 1985: 425. 

Type species: Sambirania brachysoma 


REMARKS. This genus is endemic to Madagascar. 

75. aeneicollis Descarpentries 

Sambirania aeneicollis Descarpentries, 1968: 


76. brachysoma Obenberger 

Sambirania brachysoma Obenberger, 1942: 8–9 

(orig. desc.), figs 1, 6; Descarpentries, 1968: 

929, fig. 5. 

77. dubia Descarpentries 

Sambirania dubia Descarpentries, 1968: 930 


78. impedita Descarpentries 

Sambirania impedita Descarpentries, 1968: 929 

(orig. desc.), figs 4, 8. 

79. incerta Descarpentries 

Sambirania incerta Descarpentries, 1968: 930 


80. inflata Descarpentries 

Sambirania inflata Descarpentries, 1968: 929 

(orig. desc.), figs 6, 7. 

81. plaicollis Descarpentries 

Sambirania plaicollis Descarpentries, 1968: 930 


82. rotundipennis Descarpentries 

Sambirania rotundipennis Descarpentries, 1968: 


83. suspecta Descarpentries 

Sambirania suspecta Descarpentries, 1968: 930 


Genus 25. SAMBUS Deyrolle 

Sambus Deyrolle, 1864: 210 (orig. desc.); 

Gemminger and Harold, 1869: 1434; 


1893: 119; Obenberger, 1934: 864; 


Bellamy, 1985: 425; 1988: 111; 1991: 8;


Holynski, 1993: 15. 

Type species: Sambus lafertei Deyrolle (subsequent 

designation: Toyama, 1986: 58). 

REMARKS. This genus is widespread in the Old 

World with the exception of the western Palaearctic. 

Sambus is allied to Pseudagrilus with the synapomorphy 

of saltatorial hind legs. 

84. douquetteae Descarpentries 

Sambus douquetteae Descarpentries, 1968: 928 


85. madagascariensis Descarpentries 

Sambus madagascariensis Descarpentries, 

1968: 928 (orig. desc.). 


Sambus peyrierasi Descarpentries, 1968: 927 


87. seyrigi Descarpentries 

Sambus seyrigi Descarpentries, 1968: 928 (orig. 



Sambus vadoni Descarpentries, 1968: 927 (orig. 


Genus 26. SERANAMBIA Descarpentries 

Seranambia Descarpentries, 1974: 584–585 

(orig. desc.); Bellamy, 1985: 424. 

Type species: Seranambia nodosa Descarpentries 


REMARKS. This genus is endemic to Madagascar. 

89a. nodosa Descarpentries 

Seranambia nodosa Descarpentries, 1974: 

585–586 (orig. desc.), figs 9, 10. 

89b. nodosa ssp. vondrozensis Descarpentries 

Seranambia nodosa ssp. vondrozensis 

Descarpentries, 1974: 586 (orig. desc.). 

90. succinicola Descarpentries 

Seranambia succinicola Descarpentries, 1974: 

586–587 (orig. desc.), fig. 11. 

Genus 27. SEYRIGIA Théry 

Seyrigia Théry, 1937: 104–105 (orig. desc.); 

Bellamy, 1985: 424. 

Type species: Seyrigia albopunctata Théry (original 

designation). 


and the single species seems to be involved in a 

mimicry complex with Chloricala balachowskyi. 

91. albopunctata Théry 

Seyrigia albopunctata Théry, 1937: 106–107 


Genus 28. SUAREZIA Théry 

Suarezia Théry, 1912: 119–120 (orig. desc., nom. 

praeocc. Suarezius Fairmaire, lapsus 

calami); Obenberger, 1922: 127; 1934: 795; 

Théry, 1936a: 60; Bellamy, 1985: 423. 

Type species: Suarezia albicollis Théry (original 

monotypy). 

Suarezina Théry, 1936a: 60 (nom. nov. Suarezia 

Théry, nom. superfl.); Bellamy, 1985: 423. 


and may be distantly related to the Afrotropical 

genus Euchroaria Obenberger. 

92. albicollis Théry 

Suarezia albicollis Théry, 1912: 120–121 (orig. 

desc.); Obenberger, 1934: 795. 

Undefined placement 

The following 11 species were originally 

described in one of two coraebine genera: 

Discoderes Chevrolat or Phlocteis Kerremans. 

Discoderes was last discussed by Bellamy (1986) 

and is restricted to sub-Saharan Africa, with two 

species distributed in habitats directly below the 

Sahara. Phlocteis is a small genus with three 

sub-Saharan African species also last discussed by 

Bellamy (1986). As noted above, two Madagascan 

species, P. hova Théry and P. humeralis (Waterhouse), 

are placed therein. The types of each of 

these species require study and comparison to the 

currently more restricted definition of these two 

genera and the other Madagascan coraebines 

before transfers to other genera will be possible. 

93. ambanjana (Obenberger) 

Phlocteis ambanjana Obenberger, 1944: 

421–422 (orig. desc.). 

94. ampliata (Fairmaire) 

Discoderes ampliata Fairmaire, 1905: 124 (orig. 

desc.); Théry, 1905: 178; Marie and Lesne, 

1917: 40. Phlocteis ampliata: Obenberger, 

1928: 341; 1934: 796. 

95. cavifrons (Fairmaire) 

Discoderes cavifrons Fairmaire, 1904: 234–235 

(orig. desc.); Théry, 1905: 178; Marie and 

Lesne, 1917: 40. 

Phlocteis cavifrons: Obenberger, 1928: 341; 

1934: 796.


96. goudotii (Thomson) 

Discoderes goudotii Thomson, 1878: 82–83 

(orig. desc.); Kerremans, 1885: 151; 1892: 

236; 1903: 223; Alluaud, 1900: 196; Théry, 

1905: 114; 1936a: 60. 

Discoderes goudoti (lapsus calami): Kerremans, 

1885: 151; 1892: 236; 1903: 223; Alluaud, 

1900: 196; Théry, 1905: 114; 1936a: 60. 

Phlocteis goudoti: Obenberger, 1934: 797. 

97. iridipennis (Obenberger) 

Phlocteis iridipennis Obenberger, 1944: 

420–421 (orig. desc.). 

98. mucorea (Fairmaire) 

Discoderes mucorea Fairmaire, 1902a: 80 (orig. 



Phlocteis mucorea: Obenberger, 1928: 341; 

1934: 798. 

REMARKS. Obenberger (1928) identified this 

species as one warranting a ‘new genus’ in a brief 

discussion about the genera Discoderes and 

Phlocteis. 

99. ochraceopicta (Fairmaire) 

Discoderes ochraceopicta Fairmaire, 1888: 56 

(orig. desc.); Kerremans, 1892: 237; Alluaud, 

1900: 197; Kerremans, 1903: 223; Théry, 

1905: 114, fig. 19; Marie and Lesne, 1917: 40; 

Théry, 1937: 98. 

Phlocteis ochraceopicta: Obenberger, 1934: 798. 

100. perrieri (Fairmaire) 

Discoderes perrieri Fairmaire, 1900: 244 (orig. 



Phlocteis perrieri: Obenberger, 1928: 341; 1934: 

798. 

101. polychroa (Fairmaire) 

Discoderes polychroa Fairmaire, 1904: 234 

(orig. desc.); Théry, 1905: 177; Marie and 

Lesne, 1917: 40. 

Phlocteis polychroa: Obenberger, 1928: 341; 

1934: 798. 

102. scriptus (Théry) 

Discoderes scriptus Théry, 1937: 85–86 (orig. 


103. simplicifrons (Fairmaire) 

Discoderes simplicifrons Fairmaire, 1901: 163 

(orig. desc.); Kerremans, 1903: 223; Théry, 

1905: 115; Marie and Lesne, 1917: 40. 

Phlocteis simplicifrons: Obenberger, 1934: 799. 

Doubtful taxa 

Anadora hova Kerremans, 1899: 277 (orig. 

desc.); Alluaud, 1900: 197; Kerremans, 1903: 

258; Théry, 1905: 125; 1927: 260; 

Obenberger, 1934: 794 (howa, lapsus 

calami). 

REMARKS. Anadora Kerremans is another endemic 

Subsaharan African genus last revised by Bellamy 

(1986). Théry (1905) listed this taxon as a member 

of the Madagascan fauna and later (Théry 1927) 

synonymized it with A. mechowi Quedenfeldt. 

Obenberger (1934), however, listed it as a valid 

species in his catalogue. Since there are no other 

specimens of Anadora in the vast Madagascan 

coraebine collection in the Paris museum, I support 

the synonymy and do not consider either the genus 

or species as valid members of the Madagascan 

fauna. 

In the Paris museum general collection and in the 

Obenberger collection in Prague, there are a number 

of Madagascan coraebines labelled as ‘types’ 

of various genera and species. These are taxa 

apparently intended for description by 

Descarpentries, Théry (both Paris collection), or 

Obenberger, and thus are simply manuscript or ‘in 

collection’ names. None of these names appear to 

be valid as I cannot find descriptions or any mention 

in the literature or the Zoological Record. I will not list 

any of these names here as that would add confusion 

to the discussion and nomina nuda to the literature. 

Should any of these names be found in the 

literature, they will be discussed in a forthcoming 

contribution to this series. 

ACKNOWLEDGEMENTS 

For access to the collections under their respective 

care or for the loan of specimens, I thank J. J. 

Menier, Museum National d’Histoire Naturelle, 

Paris, France; S. Bílý, Narodni Museum, Prague, 

Czech Republic; J. Cools, Institut Royal des 

Sciences Naturelles de Belgique, Brussels, 

Belgium; R. Holynski, Milanowek, Poland; M. Kerley, 

The Natural History Museum, London, United Kingdom; 

T. Neef de Sainval, Brussels; E. Sprecker, 

Naturhistorisches Museum, Basel, Switzerland and 

M. Uhlig, Museum für Naturkunde der Humboldt 

Universität, Berlin, Germany. I also thank the late A. 

Descarpentries for inviting me to join him in working 

on this great treasure of beautiful beetles back in 

1988; sorry it took so long to get started! Lastly to 

Ted MacRae, Monsanto Corporation, St Louis, 

Missouri and Rick Westcott, Oregon Department of 

Agriculture, Salem, Oregon, my appreciation for 

their reading of the original draft and for suggestions 

toward its improvement.


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NOTE ADDED IN PROOF 

A new paper by Kuban et al. (2001) was received after this paper was in proof stage. The authors recognized 

Coraebini at the tribal rank and proposed a phylogenetic arrangement of ten subtribes. The entire 

compliment of Madagascan genera were placed in the new subtribe Toxoscelina Majer. 

Reference 

Kubán, 

V., Majer, K. and Kolibá, J., 2001. Classification of the tribe Coraebini Bedel, 1921 (Coleoptera, Buprestidae, 

Agrilinae). Acta Musei Moraviae, Scientiae biologicae (Brno) 85 (2000): 185–287.

The Madagascan Coraebina Bedel, 1921 (Coleoptera: Buprestidae ...

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