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770<br />

COLLEGE ENTOMOLOGY<br />

lower ones smaller, males holoptic or dichoptic. Scutellum bare or hairy.<br />

Tibire with or without spurs; fore pair with one, two, or none; hind pair with<br />

one or two. Pulvilli present. Empodia vestigial, narrow or pad-like. Wings<br />

hyaline or with fuscous spots; venation distinct; usually five (rarely four)<br />

posterior cells; discal cell present or absent; costa extending around tip of<br />

wing; rom distinct; squamre absent or present, sometimes vestigial. Larvre<br />

variable with small, partially exserted head which is prolonged in Xylophagus;<br />

maxillary palpi well developed; mandibles long, vertical, protrusile, simple or<br />

hooked; body H-segmented and with pseudopods. Aquatic forms flattened and<br />

Sc<br />

FIG. 280. Wing of the large apiocerid fly, Raphiomidas xmzthos Townsend.<br />

with two round protrusile blood gills on the last segment below the long anal<br />

filaments. Terrestrial forms generally cylindrical; with simple or spines<br />

pseudopods on the fifth or on each abdominal segment; lateral spine-like fila"<br />

ments sometimes present; segments X and XI each with a row of hooklets in<br />

Vermileo; last segment with hairy process; spiracles present. These flies frequent<br />

wooded areas near fresh water. Aquatic forms inhabit fresh water and<br />

are predacious on small aquatic animals. Terrestrial species are likewise<br />

predaciOUS and live under bark of trees, in dung, fungi, soil, and wood. Members<br />

of the remarkable genus Vermileo excavate conical pits in the sand and<br />

dust at the bottom of which they lie concealed to capture and feed on the<br />

ants that they entrap in much the same manner as ant lions. That two widely<br />

separated groups of insects with such amazing structural differences should<br />

occupy exactly the same niche in nature is remarkable. Especially is it astonishing<br />

that such a simple, helpless-looking creature as this fly larva should<br />

simulate the accomplishments of so well-equipped, active, and formidable a<br />

gladiator as is the ant lion. The larvre of the so-called worm-lions, Vermileo,<br />

construct smaller and steeper sloping pits than those of their rivals, but they<br />

are often very numerous. In the protected side of a giant granite boulder in<br />

the Yosemite Valley, CaliL, the author counted upwards of a thousand of<br />

these contiguous pits. The Sierra worm-lion, Vermileo comstocki Wheeler,<br />

occurs in the Sierra Nevada Mountains at an altitude of 4,000 to 9,000 ft.<br />

Members of the genera Lampromyia Macquart and Vermitigris Wheeler are<br />

closely related to Vermileo, and the larvre and adults are quite similar in habits<br />

and appearances. Species of Lampromyia have been taken in most of the Span-

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