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640<br />

COLLEGE ENTOMOLOGY<br />

Family EVANIIDlE (Leach 1812) Westwood 1840 (Evan"i'i"dre, from the<br />

Latin evanidus, frail, feeble; because of their slow movements). Ensign<br />

Flies.<br />

Short, robust, somber insects frorn 4-17 mm. in length, which carry the<br />

abdomen raised into the air like an ensign from which habit the common name<br />

is derived. Antennae 13" to 14"segmented. Wings with primitive venation,<br />

i.e., many cells and broad veins; fore wings with distinct costal cell; with or<br />

without one discoidal cell; and radial cell short and broad or absent. Abdomen<br />

with long slender pedicel attached high on the upper part of the median dorsal<br />

plate above the hind coxre; gaster small. The larvre are parasitic and have been<br />

reared from eggs of cockroaches and mantids and from the larvre of xiphydrid<br />

horntails. The family consists of about 200 species widely distributed and espe"<br />

cially well represented in Australia. The important genera are Brachygaster<br />

Leach, Evania Fab., Hyptia I11iger, Hyptz"ogaster Kieffer, Semceodogaster Bradley.<br />

The members of the related families GASTERUPTIONID.tE (Ashmead<br />

1900) Handlirsch 1925, parasitic on the larvre of bees and wasps, and AULACI­<br />

DJE Schuckard 1841, parasitic on the larvre of cerambycid beetles, have the<br />

abdomen similarly attached to the propodeum.<br />

Family ICHNEUMONIDJE Leach 1817 (Ich'neu-mon'i-dre, from the Greek<br />

lxv€up.(!)JI, a small wasp that hunts spiders; described by Aristotle and<br />

Pliny; literally a tracker). German, Schlupfwespen. French, Ichneumonides.<br />

Ichneumonids, Ichneumon Flies.<br />

Minute to large slender parasitic wasps with long, many"segmented, filiform<br />

antennre; three ocelli; well-developed mouth parts; long slender legs with di"<br />

vided trochanters; conspicuous tibial spurs, strong claws, and with an empodium.<br />

Wings usually large; rarely absent or brachypterous in both sexes; welldefined<br />

venation; costal cells of fore wings eliminated by coalescence of costal<br />

and subcostal veins in most forms; median or costal cell separated from discoidal<br />

by the veinlet m-cu. Propodeum enlarged and prolonged behind the<br />

posterior coxre; often sculptured. Abdomen long and slender; cylindrical or<br />

compressed laterally; attached to lower part of propodeum; two or three times<br />

as long as the head and thorax; ovipositor variable in length up to six times the<br />

length of the body. Eggs either deposited on the outside or inserted within the<br />

body of the host; sometimes with a stalk or tube.<br />

Larvre, especially the endoparasitic forms, undergo a complex metamorphosis<br />

with four or five different stages. First stage often with long respiratory prolongation<br />

or tail which is reabsorbed; spiracular respiration appears along with<br />

the large head in the third stage. The fully developed body is 13-segmented.<br />

The larvre are attached to the exterior walls (ectoparasitic) or may hatch within<br />

or bore into the body where they live in the body cavity of the host (endopara"<br />

sitic). The larvre may enter either the larval or pupal stages of the host, and the<br />

adult ichneumonids usually emerge from the chrysalis or pupa of the host. The<br />

most important hosts consist chiefly of the larvre and pupre .of LEPIDOPTERA,<br />

Hy}4ENOPTERA, COLEOPTERA, and DIPTERA in the order named.

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